H. M. André, M. I. Noti, and P. Lebrun, The soil fauna: the other last biotic frontier, BIODIVERSITY AND CONSERVATION, vol.21, issue.1, pp.45-56, 1994.
DOI : 10.1007/BF00115332

S. J. Arnold, Morphology, Performance and Fitness, American Zoologist, vol.23, issue.2, pp.347-361, 1983.
DOI : 10.1093/icb/23.2.347

L. Barbaro and I. Van-halder, Linking bird, carabid beetle and butterfly life-history traits to habitat fragmentation in mosaic landscapes, Ecography, vol.12, issue.2, pp.321-333, 2009.
DOI : 10.1111/j.1600-0587.2008.05546.x

C. J. Barnard, Animal Behaviour: Mechanism, Development, Function, and Evolution, p.525, 2004.
DOI : 10.1007/978-1-4615-9781-0

E. Barrios, Soil biota, ecosystem services and land productivity. Ecological 526, Economics, vol.64, pp.269-285, 2007.
DOI : 10.1016/j.ecolecon.2007.03.004

J. R. Bell, A. Mead, D. J. Skirvin, K. D. Sunderland, J. S. Fenlon et al., Do functional traits improve prediction of predation rates for a disparate group of 529 aphid predators?, C. Bulletin of Entomological Research, vol.528, issue.98, pp.587-597, 2008.

L. R. Belyea and J. Lancaster, Assembly Rules within a Contingent Ecology, Oikos, vol.86, issue.3, pp.531-402, 1999.
DOI : 10.2307/3546646

J. Stadler, On the identification of the most suitable traits for plant functional trait 534 analyses, Oikos, vol.117, pp.1533-1541, 2008.

C. Bertelsmeier, G. M. Luque, A. Confais, and F. Courchamp, Ant Profiler ? a 536 database of ecological characteristics of ants (Hymenoptera: Formicidae) Myrmecological 537 News, pp.73-76, 2012.

A. Blanck, P. A. Tedesco, and N. Lamouroux, Relationships between life-history 539 strategies of European freshwater fish species and their habitat preferences, pp.843-859, 2007.

S. Bokhorst, G. K. Phoenix, J. W. Bjerke, T. V. Callaghan, F. Huyer-brugman et al., Extreme winter warming events more negatively impact small rather than large 543 soil fauna: shift in community composition explained by traits not taxa, Global Change Biology, vol.542, issue.18, pp.1152-1162, 2012.

N. Bonada, S. Dolédec, and B. Statzner, Taxonomic and biological trait differences of stream macroinvertebrate communities between mediterranean and temperate regions: implications for future climatic scenarios, Global Change Biology, vol.31, issue.8, pp.1658-1671, 2007.
DOI : 10.1038/416389a

N. Bonada, N. Prat, V. H. Resh, and B. Statzner, Developments in aquatic insect 549 biomonitoring: A comparative analysis of recent approaches, Annual Review of Entomology, vol.550, pp.495-523, 2006.

D. Bonte, L. Lens, and J. P. Maelfait, Sand dynamics in coastal dune landscapes constrain diversity and life-history characteristics of spiders, Journal of Applied Ecology, vol.2, issue.4, pp.735-747, 2006.
DOI : 10.1111/j.1365-2664.2006.01175.x

Z. Botta-dukat, Rao's quadratic entropy as a measure of functional diversity based on multiple traits, Journal of Vegetation Science, vol.95, issue.5, pp.533-540, 2005.
DOI : 10.1111/j.1654-1103.2005.tb02393.x

M. B. Bouché, Lombriciens de France. Ecologie et Systématique, p.557, 1972.

S. Buchholz, Simulated climate change in dry habitats: do spiders respond to experimental small-scale drought?, Journal of Arachnology, vol.38, issue.2, pp.280-284, 2010.
DOI : 10.1636/P09-91.1

P. A. Buxton, Animal Life in Deserts: A Study of the Fauna in Relation to the Environment, Geographical Review, vol.15, issue.1, p.560, 1923.
DOI : 10.2307/207940

J. Cavender-bares, K. H. Kozak, P. V. Fine, and S. W. Kembel, The merging of community ecology and phylogenetic biology, Ecology Letters, vol.428, issue.Suppl. 1, pp.693-715, 2009.
DOI : 10.1111/j.1461-0248.2009.01314.x

D. Chessel, A. B. Dufour, and J. Thioulouse, The ade4 Package, R News, vol.4, pp.5-10, 2004.
URL : https://hal.archives-ouvertes.fr/hal-00427558

F. Chevenet, S. Dolédec, and D. Chessel, A fuzzy coding approach for the analysis of long-term ecological data, Freshwater Biology, vol.31, issue.3, pp.295-309, 1994.
DOI : 10.1007/BF00038688

L. J. Cole, D. I. Mccracken, P. Dennis, I. S. Downie, A. L. Griffin et al., Relationships between agricultural management and 567 ecological groups of ground beetles (Coleoptera : Carabidae) on Scottish farmland, pp.323-336, 2002.

S. Cristofoli, G. Mahy, R. Kekenbosch, and K. Lambeets, Spider communities as evaluation tools for wet heathland restoration, Ecological Indicators, vol.10, issue.3, pp.773-780, 2010.
DOI : 10.1016/j.ecolind.2009.11.013

D. Lange, H. J. Lahr, J. Van-der-pol, J. J. Faber, and J. H. , Ecological vulnerability in wildlife: Application of a species-ranking method to food chains and habitats, Environmental Toxicology and Chemistry, vol.408, issue.12, pp.2875-2880, 2010.
DOI : 10.1002/etc.336

T. Decaëns, J. J. Jimenez, C. Gioia, G. J. Measey, and P. Lavelle, The values of soil 575 animals for conservation biology, European Journal of Soil Biology, pp.23-38, 2006.

T. Decaëns, P. Margerie, M. Aubert, M. Hedde, and F. Bureau, Assembly rules 577 within earthworm communities in North-Western France?A regional analysis, 2008.

T. Decaëns, P. Margerie, J. Renault, F. Bureau, M. Aubert et al., Niche 580 overlap and species assemblage dynamics in an ageing pasture gradient in north-western 581, 2011.

S. Díaz and M. Cabido, Vive la différence: plant functional diversity matters to 583 ecosystem processes Trends in Ecology & Evolution, pp.646-654, 2001.

S. Dolédec, B. Statzner, and M. Bournard, Species traits for future biomonitoring across ecoregions: patterns along a human-impacted river, Freshwater Biology, vol.14, issue.4, pp.737-758, 1999.
DOI : 10.1111/j.1365-2427.1984.tb00039.x

D. A. Driscoll and T. Weir, Beetle Responses to Habitat Fragmentation Depend on Ecological Traits, Habitat Condition, and Remnant Size, Conservation Biology, vol.4, issue.1, pp.182-194, 2005.
DOI : 10.2307/2405057

H. Ellenberg, Vegetation ecology of central, Europe, 1988.

C. R. Fonseca and G. Ganade, Species functional redundancy, random extinctions and the stability of ecosystems, Journal of Ecology, vol.81, issue.1, pp.118-125, 2001.
DOI : 10.1016/0038-0717(94)90131-7

B. Fournier, E. Samaritani, J. Shrestha, E. A. Mitchell, and R. C. Le-bayon, Patterns of earthworm communities and species traits in relation to the perturbation gradient of a restored floodplain, Applied Soil Ecology, vol.59, pp.87-95, 2012.
DOI : 10.1016/j.apsoil.2012.03.015

E. A. Frimpong and P. L. Angermeier, Trait-Based Approaches in the Analysis of 595 Stream Fish Communities, American Fisheries Society Symposium, vol.73, pp.109-136, 2010.

E. Garnier and M. L. Navas, A trait-based approach to comparative functional plant 597 ecology: concepts, methods and applications for agroecology. A review Agronomy for 598 Sustainable Development, pp.365-399, 2012.

M. Gerisch, Habitat disturbance and hydrological parameters determine the body size 600 and reproductive strategy of alluvial ground beetles, Zookeys, pp.353-370, 2011.

M. Gerisch, V. Agostinelli, K. Henle, and F. Dziock, More species, but all do the same: contrasting effects of flood disturbance on ground beetle functional and species diversity, Oikos, vol.5, issue.4, pp.508-515, 2012.
DOI : 10.1111/j.1600-0706.2011.19749.x

P. S. Giller, The diversity of soil communities, the ?poor man's tropical rainforest?, Biodiversity and Conservation, vol.5, issue.3, pp.135-168, 1996.
DOI : 10.1007/BF00055827

H. Gisin, Ökologie und Lebensgemeinschaften der Collembolen im Schweizerischen 608, 1943.

H. Gitay and I. R. Noble, What are functional types and how should we seek them?, Plant Functional Types, pp.3-19, 1997.

J. Gobat, M. Aragno, and W. Matthey, Le sol vivant, 1998.

M. Gobbi and D. Fontaneto, Biodiversity of ground beetles (Coleoptera: Carabidae) in different habitats of the Italian Po lowland, Agriculture, Ecosystems & Environment, vol.127, issue.3-4, pp.619-273, 2008.
DOI : 10.1016/j.agee.2008.04.011

P. S. Grimbacher and N. E. Stork, Seasonality of a Diverse Beetle Assemblage Inhabiting Lowland Tropical Rain Forest in Australia, Biotropica, vol.50, issue.3, pp.328-337, 2009.
DOI : 10.1111/j.1744-7429.2008.00477.x

M. Hedde, F. Van-oort, and I. Lamy, Functional traits of soil invertebrates as indicators for exposure to soil disturbance, Environmental Pollution, vol.164, pp.59-65, 2012.
DOI : 10.1016/j.envpol.2012.01.017

R. Hesse, Tiergeographie auf oekologischer Grundlage, p.625, 1924.

K. Huebner, Z. Lindo, and M. J. Lechowicz, Post-fire succession of collembolan communities in a northern hardwood forest, European Journal of Soil Biology, vol.48, pp.59-65, 2012.
DOI : 10.1016/j.ejsobi.2011.10.004

L. S. Jelaska, A. Jesovnik, S. D. Jelaska, A. Pirnat, M. Kucinic et al., 628 Variations of carabid beetle and ant assemblages, and their morpho-ecological traits within 629 natural temperate forests in Medvednica Nature Park, pp.475-486, 2010.

N. Jennings and M. J. Pocock, Relationships between Sensitivity to Agricultural 631 Intensification and Ecological Traits of Insectivorous Mammals and Arthropods, Conservation Biology, vol.632, issue.23, pp.1195-1203, 2009.

J. J. Jimenez, T. Decaëns, and J. P. Rossi, Soil environmental heterogeneity allows 634 spatial co-occurrence of competitor earthworm species in a gallery forest of the Colombian 635 'Llanos'. Oikos, pp.915-926, 2012.

M. B. Jones, M. P. Schildhauer, O. J. Reichman, and S. Bowers, The new 637 bioinformatics: Integrating ecological data from the gene to the biosphere Palo Alto, Annual Review of 638 Ecology Evolution and Systematics, pp.519-544, 2006.

M. Karen, J. O-'halloran, J. Breen, P. Giller, J. Pithon et al., Distribution and composition of carabid beetle (Coleoptera, Carabidae) communities across the plantation forest cycle???Implications for management, Forest Ecology and Management, vol.256, issue.4, pp.624-632, 2008.
DOI : 10.1016/j.foreco.2008.05.005

P. A. Keddy, Assembly and response rules: two goals for predictive community ecology, Journal of Vegetation Science, vol.119, issue.2, pp.157-164, 1992.
DOI : 10.2307/3235676
URL : http://citeseerx.ist.psu.edu/viewdoc/summary?doi=10.1.1.668.5900

M. G. Kibblewhite, K. Ritz, and M. J. Swift, Soil health in agricultural systems, Philosophical Transactions of the Royal Society B: Biological Sciences, vol.304, issue.5677, pp.685-701, 2008.
DOI : 10.1126/science.1097394

K. Lambeets, M. L. Vandegehuchte, J. P. Maelfait, and D. Bonte, Understanding the impact of flooding on trait-displacements and shifts in assemblage structure of predatory arthropods on river banks, Journal of Animal Ecology, vol.9, issue.Suppl. S4, pp.1162-1174, 2008.
DOI : 10.1111/j.1365-2656.2008.01443.x

K. Lambeets, M. L. Vandegehuchte, J. P. Maelfait, and D. Bonte, Integrating environmental conditions and functional life-history traits for riparian arthropod conservation planning, Biological Conservation, vol.142, issue.3, pp.625-637, 2009.
DOI : 10.1016/j.biocon.2008.11.015

P. R. Langlands, K. E. Brennan, V. W. Framenau, and B. Y. Main, Predicting the post-fire responses of animal assemblages: testing a trait-based approach using spiders, Journal of Animal Ecology, vol.43, issue.3, pp.558-568, 2011.
DOI : 10.1111/j.1365-2656.2010.01795.x

M. Laporte, I. Mougenot, and E. Garnier, ThesauForm???Traits: A web based collaborative tool to develop a thesaurus for plant functional diversity research, Ecological Informatics, vol.11, 2012.
DOI : 10.1016/j.ecoinf.2012.04.004

P. Lavelle, T. Decaëns, M. Aubert, S. Barot, M. Blouin et al., Soil invertebrates and??ecosystem services, European Journal of Soil Biology, vol.42, issue.42, pp.3-15, 2006.
DOI : 10.1016/j.ejsobi.2006.10.002

P. Lavelle and A. V. Spain, Soil ecology, p.662, 2001.
DOI : 10.1007/978-94-017-5279-4
URL : https://hal.archives-ouvertes.fr/bioemco-00455666

N. Harguindeguy, C. Roumet, C. Urcelay, J. G. Canadell, D. E. Pataki et al., Plant functional types: are we getting any closer to the Holy Grail? Terrestrial 665 ecosystems in a changing world, pp.149-164, 2007.

S. Lavorel and E. Garnier, Predicting changes in community composition and ecosystem functioning from plant traits: revisiting the Holy Grail, Functional Ecology, vol.5, issue.5, pp.545-556, 2002.
DOI : 10.1016/0169-5347(92)90006-W

S. Lavorel, S. Mcintyre, J. Landsberg, and T. D. Forbes, Plant functional 670 classifications: from general groups to specific groups based on response to disturbance. 671 Trends in Ecology &amp, Evolution, vol.12, pp.474-478, 1997.

L. Viol, I. Julliard, R. Kerbiriou, C. De-redon, L. Carnino et al., Plant and spider communities benefit differently from the presence of planted 674 hedgerows in highway verges, Biological Conservation, vol.141, pp.673-1581, 2008.

N. Lindberg and J. Bengtsson, Population responses of oribatid mites and collembolans after drought, Applied Soil Ecology, vol.28, issue.2, pp.163-174, 2005.
DOI : 10.1016/j.apsoil.2004.07.003

J. Madin, S. Bowers, M. Schildhauer, S. Krivov, D. Pennington et al., An 681 ontology for describing and synthesizing ecological observation data, Ecological Informatics, vol.682, issue.2, pp.279-296, 2007.

M. Makkonen, M. P. Berg, J. R. Van-hal, T. V. Callaghan, M. C. Press et al., 684 Traits explain the responses of a sub-arctic Collembola community to climate manipulation, Soil Biology & Biochemistry, vol.685, issue.43, pp.377-384, 2011.

A. Malmstrom, Life-history traits predict recovery patterns in Collembola species after fire: A 10 year study, Applied Soil Ecology, vol.56, pp.35-42, 2012.
DOI : 10.1016/j.apsoil.2012.02.007

B. J. Mcgill, B. J. Enquist, E. Weiher, and M. Westoby, Rebuilding community ecology from functional traits, Trends in Ecology & Evolution, vol.21, issue.4, pp.178-185, 2006.
DOI : 10.1016/j.tree.2006.02.002

W. K. Michener, Quantitatively Evaluating Restoration Experiments: Research Design, Statistical Analysis, and Data Management Considerations, Restoration Ecology, vol.5, issue.4, pp.324-337, 1997.
DOI : 10.1038/335154a0

W. K. Michener, Meta-information concepts for ecological data management, 694 Ecological Informatics, pp.3-7, 2006.
DOI : 10.1016/j.ecoinf.2005.08.004

A. T. Moles, D. I. Warton, L. Warman, N. G. Swenson, S. W. Laffan et al., Global patterns in plant height, Journal of Ecology, vol.14, issue.5, pp.923-932, 2009.
DOI : 10.1111/j.1365-2745.2009.01526.x

M. Moretti and C. Legg, Combining plant and animal traits to assess community functional responses to disturbance, Ecography, vol.428, issue.2, pp.299-309, 2009.
DOI : 10.1111/j.1600-0587.2008.05524.x

S. Nylin and K. Gotthard, Plasticity in Life-History Traits. Annual Review of 701 Entomology, pp.63-83, 1998.

A. S. Pearse, Animal Ecology, 1926.

D. Piron, A. Richard, A. Bispo, C. Grand, L. Galsomies et al., Earthworm 705 indicators as tools for soil monitoring, characterization and risk assessment. An example from 706 the national Bioindicator programme (France), Pedobiologia, pp.77-87, 2011.

N. L. Poff, J. D. Olden, N. K. Vieira, D. S. Finn, M. P. Simmons et al., Functional trait niches of North American lotic insects: traits-based ecological applications in light of phylogenetic relationships, Journal of the North American Benthology 711 Society, pp.730-755, 2006.
DOI : 10.1899/0887-3593(2006)025[0730:FTNONA]2.0.CO;2

J. F. Ponge, F. Dubs, S. Gillet, J. P. Sousa, and P. Lavelle, Decreased biodiversity in soil springtail communities: the importance of dispersal and landuse history in heterogeneous landscapes, Soil Biology and Biochemistry, vol.38, issue.5, pp.1158-1161, 2006.
DOI : 10.1016/j.soilbio.2005.09.004
URL : https://hal.archives-ouvertes.fr/hal-00495402

J. H. Porter, Scientific databases for environmental research Data and information management in the ecological sciences: A 717 resource guide R: A Language and Environment for Statistical 720 Computing, 718 Long Term Ecological Research Network Office. 719 R Development Core Team, pp.41-46, 1998.

R. Relya and A. , Morphological and behavioral plasticity of larval anurans in response to 722 different predators, Ecology Letters, vol.82, p.723, 2001.

I. Ribera, S. Doledec, I. S. Downie, and G. N. Foster, EFFECT OF LAND DISTURBANCE AND STRESS ON SPECIES TRAITS OF GROUND BEETLE ASSEMBLAGES, Ecology, vol.82, issue.4, pp.1112-1129, 2001.
DOI : 10.1098/rspb.1999.0696

R. Schmelz and W. E. Xylander, The Edaphobase GBIF project of germany -a new 727 online soil organism data warehouse. 16th International Colloquium on Soil Zoology, p.34, 2012.

S. Salmon and J. F. Ponge, Species traits and habitats in springtail communities: A 730 regional scale study. Pedobiologia, pp.295-301, 2012.

H. Schmallfuss, Eco-morphological strategies in terrestrial isopods Biology (The) of 732 terrestrial isopods, Symposium, vol.53, pp.49-63, 1983.

K. Semper, Die Natürlichen Existenzbedingungen der Thiere, pp.734-735, 1880.

V. E. Shelford, Animal Communities in Temperate America, 1913.
DOI : 10.5962/bhl.title.21631

P. M. Silva, C. A. Aguiar, I. D. Faria-e-silva, and A. R. Serrano, Orchard and riparian habitats enhance ground dwelling beetle diversity in Mediterranean agro-forestry systems, Biodiversity and Conservation, vol.134, issue.4, pp.861-872, 2011.
DOI : 10.1007/s10531-010-9987-6

T. R. Southwood, Habitat, the Templet for Ecological Strategies?, The Journal of Animal Ecology, vol.46, issue.2, pp.336-365, 1977.
DOI : 10.2307/3817

B. Statzner, S. Dolédec, and B. Hugueny, Biological trait composition of European stream invertebrate communities: assessing the effects of various trait filter types, Ecography, vol.65, issue.Suppl. 1, pp.470-488, 2004.
DOI : 10.1111/j.0906-7590.2004.03836.x

B. Statzner, A. G. Hildrew, and V. H. Resh, Species traits and environmental 745 constraints: Entomological research and the history of ecological theory, Annual Review, pp.746-291, 2001.

S. C. Stearns, The Evolution of Life Histories, p.748, 1992.

M. J. Swift, O. W. Heal, and J. M. Anderson, Decomposition in Terrestrial Ecosystems, p.750, 1979.

W. Thuiller, S. Lavorel, G. Midgley, S. B. Lavergne, and T. Rebelo, Relating plant 751 traits and species distributions along bioclimatic gradients for 88 Leucadendron taxa, Ecology, vol.85, pp.752-1688, 2004.

R. Tropek, L. Spitzer, and M. Konvicka, Two groups of epigeic arthropods differ in colonising of piedmont quarries: the necessity of multi-taxa and life-history traits approaches in the monitoring studies, Community Ecology, vol.9, issue.2, pp.177-184, 2008.
DOI : 10.1556/ComEc.9.2008.2.6

A. J. Vanbergen, B. A. Woodcock, M. Koivula, J. Niemelä, D. J. Kotze et al., Trophic level 759 modulates carabid beetle responses to habitat and landscape structure: a pan-European study, Ecological Entomology, vol.35, pp.760-226, 2010.

R. Verhagen, R. V. Diggelen, and R. Vermeulen, Community, p.767, 2008.

S. Villéger, N. W. Mason, and D. Mouillot, NEW MULTIDIMENSIONAL FUNCTIONAL DIVERSITY INDICES FOR A MULTIFACETED FRAMEWORK IN FUNCTIONAL ECOLOGY, Ecology, vol.89, issue.8, pp.2290-2301, 2008.
DOI : 10.1111/j.1461-0248.2005.00850.x

C. Violle, M. L. Navas, D. Vile, E. Kazakou, C. Fortunel et al., Let the concept of trait be functional! Oikos, pp.771-882, 2007.

G. D. Warnaffe and M. Dufrene, To what extent can management variables explain species assemblages? A study of carabid beetles in forests, Ecography, vol.69, issue.4, pp.701-714, 2004.
DOI : 10.1111/j.0906-7590.2004.03880.x

C. T. Webb, J. A. Hoeting, G. M. Ames, M. I. Pyne, L. Poff et al., A structured 775 and dynamic framework to advance traits-based theory and prediction in ecology, Ecology, vol.776, issue.13, pp.267-283, 2010.

V. Wolters, Biodiversity of soil animals and its function, European Journal of Soil Biology, vol.37, issue.4, pp.221-227, 2001.
DOI : 10.1016/S1164-5563(01)01088-3

J. P. Wright, S. Naeem, A. Hector, C. Lehman, P. B. Reich et al., Conventional functional classification schemes underestimate the relationship with 781 ecosystem functioning, Ecology Letters, vol.9, pp.780-111, 2006.

M. B. Bouché, Lombriciens de France. Ecologie et Systématique, p.119, 1972.

P. Brandmayr, Studio ecologico sui Coleotteri Carabid di due formzioni vegetali dell'Istria 120 Montana, Boll. Zool, vol.39, p.121, 1972.

F. Chevenet, S. Dolédec, and D. Chessel, A fuzzy coding approach for the analysis of long-term ecological data, Freshwater Biology, vol.31, issue.3, pp.295-309, 1994.
DOI : 10.1007/BF00038688

M. J. Davies, The contents of the crops of some British Carabid Beetles, Ent. mon. Mag, vol.95, p.124, 1953.

I. Ribera, S. Doledec, I. S. Downie, and G. N. Foster, EFFECT OF LAND DISTURBANCE AND STRESS ON SPECIES TRAITS OF GROUND BEETLE ASSEMBLAGES, Ecology, vol.82, issue.4, pp.1112-1129, 2001.
DOI : 10.1098/rspb.1999.0696