J. L. Brunner and R. S. Ostfeld, MULTIPLE CAUSES OF VARIABLE TICK BURDENS ON SMALL-MAMMAL HOSTS, Ecology, vol.22, issue.8, pp.2259-2272, 2008.
DOI : 10.1073/pnas.94.1.338

A. Butet, G. Paillat, and Y. Delettre, Seasonal changes in small mammal assemblages from field boundaries in an agricultural landscape of western France, Agriculture, Ecosystems & Environment, vol.113, issue.1-4, pp.364-369, 2006.
DOI : 10.1016/j.agee.2005.10.008
URL : https://hal.archives-ouvertes.fr/hal-00019506

A. Chastagner, M. Moinet, G. Perez, E. Roy, K. D. Mccoy et al., Prevalence of Anaplasma phagocytophilum in small rodents in France, Ticks and Tick-borne Diseases, vol.7, issue.5, pp.988-991, 2016.
DOI : 10.1016/j.ttbdis.2016.05.005
URL : https://hal.archives-ouvertes.fr/hal-01328852

B. Close, K. Banister, V. Baumans, E. M. Bernoth, N. Bromage et al., Recommendations for euthanasia of experimental animals: Part 2, Laboratory Animals, vol.31, issue.1, pp.1-32, 1997.
DOI : 10.1258/002367797780600297

M. J. Cotton and C. H. Watts, The ecology of the tick Ixodes trianguliceps Birula (Arachnida; Acarina; Ixodoidea), Parasitology, vol.13, issue.03, pp.525-531, 1967.
DOI : 10.1017/S0031182000072401

J. W. Courtney, L. M. Kostelnik, N. S. Zeidner, and R. F. Massung, Multiplex Real-Time PCR for Detection of Anaplasma phagocytophilum and Borrelia burgdorferi, Journal of Clinical Microbiology, vol.42, issue.7, pp.3164-3168, 2004.
DOI : 10.1128/JCM.42.7.3164-3168.2004

L. Crespin, R. Verhagen, N. C. Stenseth, N. G. Yoccoz, and A. Pre, Survival in fluctuating bank vole populations: seasonal and yearly variations, Oikos, vol.40, issue.3, pp.467-479, 2002.
DOI : 10.1080/02664760120108764

B. Cull, A. G. Vaux, L. J. Ottowell, E. L. Gillingham, and J. M. Medlock, Tick infestation of small mammals in an English woodland, Journal of Vector Ecology, vol.2, issue.pt 2, pp.74-83, 2017.
DOI : 10.1016/j.ijppaw.2012.11.003

T. A. Dallas, S. A. Fore, K. , and H. , burden, Journal of Vector Ecology, vol.41, issue.2, pp.338-341, 2012.
DOI : 10.2307/1377530

M. Daniel, M. Malý, V. Danielová, K. Bohumír, and P. Nuttall, Abiotic predictors and annual seasonal dynamics of Ixodes ricinus, the major disease vector of Central Europe, Parasites & Vectors, vol.18, issue.Suppl 1, pp.1-12, 2015.
DOI : 10.1016/j.ijmm.2008.01.010

G. Devevey and D. Brisson, The effect of spatial heterogenity on the aggregation of ticks on white-footed mice, Parasitology, vol.36, issue.07, pp.915-940, 2012.
DOI : 10.1017/S0031182000046382

A. Dizij and K. Kurtenbach, Clethrionomys glareolus, but not Apodemus flavicollis, acquires resistance to lxodes ricinus L, the main European vector of Borrelia burgdorferi, Parasite Immunology, vol.30, issue.4, pp.177-183, 1995.
DOI : 10.2307/3272160

A. D. Dobson, J. L. Taylor, R. , and S. E. , Tick (Ixodes ricinus) abundance and seasonality at recreational sites in the UK: Hazards in relation to fine-scale habitat types revealed by complementary sampling methods, Ticks and Tick-borne Diseases, vol.2, issue.2, pp.67-74, 2011.
DOI : 10.1016/j.ttbdis.2011.03.002

J. M. Doby, G. Bigaignon, and B. Degeilh, Importance potentielle comparée du mulot gris (Apodemus sylvaticus) et du campagnol roussâtre (Clethrionomys glareolus) dans l'épidémiologie de la borréliose de Lyme en milieu forestier dans l'ouest de la France, pp.271-293, 1992.

J. M. Doby, G. Bigaignon, H. Launay, C. Costil, and O. Lorvellec, Presence of Borrelia burgdorferi, agent of tick spirochaetosis, Ixodes (Exopalpiger) trianguliceps Birula, 1895 and Ixodes (Ixodes) acuminatus NeumannAcari: Ixodidae) and in <i>Ctenophthalmus bae, 1901.

L. Ferreri, M. Giacobini, P. Bajardi, L. Bertolotti, L. Bolzoni et al., Pattern of Tick Aggregation on Mice: Larger Than Expected Distribution Tail Enhances the Spread of Tick-Borne Pathogens, PLoS Computational Biology, vol.48, issue.11, p.1003931, 2014.
DOI : 10.1371/journal.pcbi.1003931.s002

J. Franke, A. Hildebrandt, and W. Dorn, Exploring gaps in our knowledge on Lyme borreliosis spirochaetes ??? Updates on complex heterogeneity, ecology, and pathogenicity, Ticks and Tick-borne Diseases, vol.4, issue.1-2, pp.11-25, 2013.
DOI : 10.1016/j.ttbdis.2012.06.007

L. Gern, E. Rouvinez, L. N. Toutoungi, and E. Godfroid, Transmission cycles of Borrelia burgdorferi sens lato involving Ixodes ricinus and/or I. hexagonus ticks and the European hedgehog, Erinaceus europaeus, in suburban and urban areas in Switzerland, Folia Parasitol, vol.44, pp.309-314, 1997.

L. Gern, M. Siegenthaler, C. M. Hu, S. Leuba-garcia, P. F. Humair et al., Borrelia burgdorferi in rodents (Apodemus flavicollis andA. sylvaticus): Duration and enhancement of infectivity forIxodes ricinus ticks, European Journal of Epidemiology, vol.79, issue.1, pp.75-80, 1994.
DOI : 10.1093/jmedent/30.1.54

B. Gilot, J. C. Beaucournu, G. Pautou, A. Fayard, and E. Moncada, Contribution à la connaissance de la répartition et de l'écologie d'Ixodes trianguliceps en France, et plus particulièrement dans le Sud-Est, Acta Trop, vol.33, pp.254-286, 1976.

B. Godsall, T. Coulson, and A. F. Malo, From physiology to space use: energy reserves and androgenization explain home-range size variation in a woodland rodent, Journal of Animal Ecology, vol.30, issue.1, pp.126-135, 2013.
DOI : 10.1007/BF00166702

B. J. Goodwin, R. S. Ostfeld, and E. M. Schauber, Spatiotemporal Variation in a Lyme Disease Host and Vector: Black-Legged Ticks on White-Footed Mice, Vector-Borne and Zoonotic Diseases, vol.1, issue.2, pp.129-167, 2001.
DOI : 10.1089/153036601316977732

J. S. Gray and G. Lohan, The development of a sampling method for the tick Ixodes ricinus and its use in a redwater fever area, Annals of Applied Biology, vol.54, issue.3, pp.421-427, 1982.
DOI : 10.1071/ZO9610752

A. Harrison, M. Scantlebury, M. , and W. I. , Body mass and sex-biased parasitism in wood mice Apodemus sylvaticus, Oikos, vol.15, issue.7, pp.1099-1104, 2010.
DOI : 10.1093/jmedent/13.4-5.389

D. Heylen, E. Tijsse, M. Fonville, E. Matthysen, and H. Sprong, s.l. in a bird tick community, Environmental Microbiology, vol.11, issue.2, pp.663-673, 2013.
DOI : 10.1089/vbz.2011.0653

T. R. Hofmeester, E. C. Coipan, S. E. Van-wieren, H. H. Prins, W. Takken et al., Few vertebrate species dominate the Borrelia burgdorferi s.l. life cycle, Environ. Res. Lett, vol.11, p.16, 2016.

M. J. Hubbard, A. S. Baker, C. , and K. J. , Distribution of Borrelia burgdorferi s.l. spirochaete DNA in British ticks (Argasidae and Ixodidae) since the 19th Century, assessed by PCR, Medical and Veterinary Entomology, vol.12, issue.1, 1998.
DOI : 10.1046/j.1365-2915.1998.00088.x

V. L. Hughes and S. E. Randolph, TESTOSTERONE DEPRESSES INNATE AND ACQUIRED RESISTANCE TO TICKS IN NATURAL RODENT HOSTS: A FORCE FOR AGGREGATED DISTRIBUTIONS OF PARASITES, Journal of Parasitology, vol.65, issue.1, pp.49-54, 2001.
DOI : 10.1645/0022-3395(2001)087[0049:TDIAAR]2.0.CO;2

V. L. Hughes and S. E. Randolph, Testosterone increases the transmission potential of tickborne parasites, Parasitology, vol.123, pp.365-371, 2001.

P. F. Humair, O. Rais, and L. Gern, Transmission of Borrelia afzelii from Apodemus mice and Clethrionomys voles to Ixodes ricinus ticks: differential transmission pattern and overwintering maintenance, Parasitology, vol.118, issue.1, pp.33-42, 1999.
DOI : 10.1017/S0031182098003564

P. F. Humair, N. Turrian, A. Aeschilimann, and L. Gern, Borrelia burgdorferi in a focus of Lyme borreliosis: epizootiologic contribution of small mammals, Folia Parasitol, vol.40, pp.65-70, 1993.

M. Jacquot, D. Abrial, P. Gasqui, S. Bord, M. Marsot et al., Multiple independent transmission cycles of a tick-borne pathogen within a local host community, Scientific Reports, vol.19, issue.1, pp.1-12, 2016.
DOI : 10.1103/PhysRevE.70.066111
URL : https://hal.archives-ouvertes.fr/hal-01365425

S. Jahfari, E. C. Coipan, M. Fonville, A. D. Van-leeuwen, P. Hengeveld et al., Circulation of four Anaplasma phagocytophilum ecotypes in Europe, Parasites & Vectors, vol.7, issue.1, p.365, 2014.
DOI : 10.1186/1756-3305-7-365

F. Jongejan and G. Uilenberg, The global importance of ticks, Parasitology, vol.96, issue.07, p.3, 2005.
DOI : 10.1016/S0169-4758(99)01472-6

F. Jouda, J. Perret, and L. Gern, Sensu Lato in Switzerland: Spatio-Temporal Pattern at a Regional Scale, Vector-Borne and Zoonotic Diseases, vol.4, issue.1, pp.23-32, 2004.
DOI : 10.1089/153036604773082960

E. R. Kallio, M. Begon, R. J. Birtles, K. J. Bown, E. Koskela et al., in Rodents in Finland, Vector-Borne and Zoonotic Diseases, vol.14, issue.6, pp.389-93, 2014.
DOI : 10.1089/vbz.2013.1383

A. F. Katelina, Y. A. Myasnikov, and T. V. Panina, Burrow Tick Ixodes Trianguliceps in Natural Foci of Haemorrhagic Fever with Renal Syndrome in Broad-Leaved Forests of Eastern Europe, Proceedings of the 3rd International Congress of Acarology, pp.603-607, 1971.
DOI : 10.1007/978-94-010-2709-0_117

F. Keesing, J. Brunner, S. Duerr, M. Killilea, K. Logiudice et al., Hosts as ecological traps for the vector of Lyme disease, Proceedings of the Royal Society B: Biological Sciences, vol.18, issue.6, pp.3911-3919, 2009.
DOI : 10.1371/journal.pone.0002488

T. Kikkawa, Movement, Activity and Distribution of the Small Rodents Clethrionomys glareolus and Apodemus sylvaticus in Woodland, The Journal of Animal Ecology, vol.33, issue.2, pp.259-299, 1964.
DOI : 10.2307/2631

V. Kjelland, S. Stuen, T. Skarpaas, and A. Slettan, Borrelia burgdorferi sensu lato Detected in Skin of Norwegian Mountain Harres (Lepus timidus) Without Signs of Dissemination, J, 2011.

H. Korn, Changes in home range size during growth and maturation of the wood mouse (Apodemus sylvaticus) and the bank vole (Clethrionomys glareolus), Oecologia, vol.18, issue.4, pp.623-628, 1986.
DOI : 10.1007/BF00378782

K. Kurtenbach, A. Dizij, H. M. Seitz, G. Margos, S. E. Moter et al., Differential immune responses to Borrelia burgdorferi in European wild rodent species influence spirochete transmission to Ixodes ricinus L. (Acari: Ixodidae), Infect. Immun, vol.62, pp.5344-5352, 1994.

K. Kurtenbach, K. Hanincová, J. I. Tsao, G. Margos, D. Fish et al., Fundamental processes in the evolutionary ecology of Lyme borreliosis, Nature Reviews Microbiology, vol.5, issue.9, pp.660-669, 2006.
DOI : 10.1093/jmedent/22.4.408

K. Kurtenbach, H. Kampen, A. Dizij, S. Arndt, H. M. Seitz et al., Infestation of Rodents with Larval Ixodes ricinus (Acari; Ixodidae) Is an Important Factor in the Transmission Cycle of Borrelia burgdorferi s.l. in German Woodlands, Journal of Medical Entomology, vol.32, issue.6, pp.807-817, 1995.
DOI : 10.1093/jmedent/32.6.807

K. Kurtenbach, S. De-michelis, S. Etti, S. M. Schäfer, H. Sewell et al., Host association of Borrelia burgdorferi sensu lato ??? the key role of host complement, Trends in Microbiology, vol.10, issue.2, pp.74-83, 2002.
DOI : 10.1016/S0966-842X(01)02298-3

L. Hostis, M. Dumon, H. Fusade, A. Lazareff, S. Gorenflot et al., Seasonal incidence of Ixodes ricinus ticks (Acari: ixodidae) on rodents in western France, Experimental and Applied Acarology, vol.36, issue.7, pp.359-368, 1996.
DOI : 10.1093/jmedent/32.6.807

E. F. Lambin, A. Tran, S. O. Vanwambeke, C. Linard, and V. Soti, Pathogenic landscapes: Interactions between land, people, disease vectors, and their animal hosts, International Journal of Health Geographics, vol.9, issue.1, p.54, 2010.
DOI : 10.1186/1476-072X-9-54
URL : https://ij-healthgeographics.biomedcentral.com/track/pdf/10.1186/1476-072X-9-54?site=ij-healthgeographics.biomedcentral.com

X. Lambin, V. Bretagnolle, Y. , and N. G. , Vole population cycles in northern and southern Europe: Is there a need for different explanations for single pattern?, Journal of Animal Ecology, vol.70, issue.2, pp.340-349, 2006.
DOI : 10.1111/j.1365-2338.1977.tb02721.x
URL : https://hal.archives-ouvertes.fr/hal-00184655

A. J. Macdonald, D. W. Hyon, J. B. Iii, K. E. Connor, A. Swei et al., Lyme disease risk in southern California: abiotic and environmental drivers of Ixodes pacificus (Acari: Ixodidae) density and infection prevalence with Borrelia burgdorferi, Parasites & Vectors, vol.87, issue.6, pp.1-16, 2017.
DOI : 10.1007/s10584-007-9377-6

A. W. Montgomery, Population Regulation in the Wood Mouse, Apodemus sylvaticus. I. Density Dependence in the Annual Cycle of Abundance, The Journal of Animal Ecology, vol.58, issue.2, pp.465-475, 1989.
DOI : 10.2307/4842

A. Mysterud, R. Byrkjeland, L. Qviller, and H. Viljugrein, The generalist tick Ixodes ricinus and the specialist tick Ixodes trianguliceps on shrews and rodents in a northern forest ecosystem??? a role of body size even among small hosts, Parasites & Vectors, vol.53, issue.S1, p.639, 2015.
DOI : 10.1007/s10493-010-9378-4

R. S. Ostfeld, C. D. Canham, K. Oggenfuss, R. J. Winchcombe, and F. Keesing, Climate, Deer, Rodents, and Acorns as Determinants of Variation in Lyme-Disease Risk, PLoS Biology, vol.60, issue.6, p.145, 2006.
DOI : 10.1371/journal.pbio.0040145.t006

C. Pérez-eid, Les tiques : Identification, biologie, importance médicale et vétérinaire, 2007.

G. Perez, S. Bastian, A. Agoulon, A. Bouju-albert, A. Durand et al., Effect of landscape features on the relationship between Ixodes ricinus ticks and their small mammal hosts, Parasites & Vectors, vol.20, issue.August, pp.1-18, 2016.
DOI : 10.1007/s10980-004-3318-9
URL : https://hal.archives-ouvertes.fr/hal-01259231

S. E. Perkins, I. M. Cattadori, V. Tagliapietra, A. P. Rizzoli, H. et al., Empirical evidence for key hosts in persistence of a tick-borne disease, International Journal for Parasitology, vol.33, issue.9, pp.909-917, 2003.
DOI : 10.1016/S0020-7519(03)00128-0

J. Perret, O. Rais, and L. Gern, Influence of Climate on the Proportion of <I>Ixodes ricinus</I> Nymphs and Adults Questing in a Tick Population, Journal of Medical Entomology, vol.41, issue.3, pp.361-365, 2004.
DOI : 10.1603/0022-2585-41.3.361

B. Pichon, D. Egan, M. Rogers, and J. Gray, L. (Acari: Ixodidae), Journal of Medical Entomology, vol.40, issue.5, pp.723-731, 2003.
DOI : 10.1603/0022-2585-40.5.723
URL : https://hal.archives-ouvertes.fr/hal-00457745

B. Pisanu, J. L. Chapuis, A. Dozières, F. Basset, V. Poux et al., High prevalence of Borrelia burgdorferi s.l. in the European red squirrel Sciurus vulgaris in France, Ticks and Tick-borne Diseases, vol.5, issue.1, pp.1-6, 2014.
DOI : 10.1016/j.ttbdis.2013.07.007

B. Pisanu, M. Marsot, J. Marmet, J. Chapuis, D. Reale et al., Introduced Siberian chipmunks are more heavily infested by ixodid ticks than are native bank voles in a suburban forest in France, International Journal for Parasitology, vol.40, issue.11, pp.1277-1283, 2010.
DOI : 10.1016/j.ijpara.2010.03.012

S. E. Randolph, Density-dependent acquired resistance to ticks in natural hosts, independent of concurrent infection with Babesia microti, Parasitology, vol.93, issue.04, pp.413-419, 1994.
DOI : 10.1007/BF01200449

S. E. Randolph, Seasonal Dynamics of a Host-Parasite System: Ixodes trianguliceps (Acarina: Ixodidae) and its Small Mammal Hosts, The Journal of Animal Ecology, vol.44, issue.2, pp.425-449, 1975.
DOI : 10.2307/3605

C. Rauter and T. Hartung, Prevalence of Borrelia burgdorferi Sensu Lato Genospecies in Ixodes ricinus Ticks in Europe: a Metaanalysis, Applied and Environmental Microbiology, vol.71, issue.11, pp.7203-7216, 2005.
DOI : 10.1128/AEM.71.11.7203-7216.2005

A. R. Renwick and X. Lambin, Host???parasite interactions in a fragmented landscape, International Journal for Parasitology, vol.43, issue.1, pp.27-35, 2012.
DOI : 10.1016/j.ijpara.2012.10.012

D. Richter, A. Debski, Z. Hubalek, and F. Matuschka, Ticks, Vector-Borne and Zoonotic Diseases, vol.12, issue.1, pp.21-27, 2012.
DOI : 10.1089/vbz.2011.0668

D. Richter, D. B. Schlee, R. Allgo, and F. Matuschka, Relationships of a Novel Lyme Disease Spirochete, Borrelia spielmani sp. nov., with Its Hosts in Central Europe, Applied and Environmental Microbiology, vol.70, issue.11, 2004.
DOI : 10.1128/AEM.70.11.6414-6419.2004

B. Roche, P. Rohani, A. P. Dobson, and J. Guégan, The Impact of Community Organization on Vector-Borne Pathogens, The American Naturalist, vol.181, issue.1, pp.1-11, 2013.
DOI : 10.1086/668591

L. Rollend, D. Fish, and J. E. Childs, Transovarial transmission of Borrelia spirochetes by Ixodes scapularis: A summary of the literature and recent observations, Ticks and Tick-borne Diseases, vol.4, issue.1-2, pp.46-51, 2013.
DOI : 10.1016/j.ttbdis.2012.06.008

R. Rosà and A. Pugliese, Effects of tick population dynamics and host densities on the persistence of tick-borne infections, Mathematical Biosciences, vol.208, issue.1, pp.216-240, 2007.
DOI : 10.1016/j.mbs.2006.10.002

R. Rosà, A. Pugliese, M. Ghosh, S. E. Perkins, and A. Rizzoli, in Relation to Local Climate and Host Dynamics, Vector-Borne and Zoonotic Diseases, vol.7, issue.3, pp.285-295, 2007.
DOI : 10.1089/vbz.2006.0607

E. C. Rynkiewicz, H. Hawlena, L. Durden, M. W. Hastriter, G. E. Demas et al., Associations between innate immune function and ectoparasites in wild rodent hosts, Parasitology Research, vol.23, issue.4, 2013.
DOI : 10.1111/j.1365-2435.2009.01572.x

E. Sinski, A. Pawelczyk, A. Bajer, and J. M. Behnke, Abundance of wild rodents, ticks and Set of the 95% most supported models according to their AIC weights. Models are ranked based on their AICc values; the best model is the one with the lowest AICc value. Sampling site is included as random factor. The fixed variable coefficients are given with the following baseline factors: "wood mouse" for species; "female" for sex; "2012" for year; "autumn" for season. Significance of the variables in a given model are indicated by "*" for p < 0.05R²marg": marginal pseudo- R²; "R²cond": conditional pseudo-R²; "df": degree of freedomAICc": Akaike Information criteron corrected for finite sample size, 2006.