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funding acquisition: J. Dav. and S.G.; biological data acquisition and management: C.B. and V.V.; sequence data acquisition, Acknowledgments: Analyses were performed on the computing cluster of the Montpellier Bioinformatics Biodiversity (MBB) platform, the South Green Bioinformatics platform, and the UPPMAX platform in Uppsala. Funding: This work was supported by the French National Research Agency ,
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