A. Katzourakis and R. J. Gifford, Endogenous viral elements in animal genomes, PLoS Genet, vol.6, 2010.

G. Magiorkinis, D. Blanco-melo, and R. Belshaw, The decline of human endogenous retroviruses: extinction and survival, Retrovirology, vol.12, p.25640971, 2015.

W. E. Johnson, Origins and evolutionary consequences of ancient endogenous retroviruses, Nat Rev Microbiol, vol.17, pp.355-370, 2019.

C. Feschotte and C. Gilbert, Endogenous viruses: insights into viral evolution and impact on host biology, Nat Rev Genet, vol.13, pp.283-296, 2012.
URL : https://hal.archives-ouvertes.fr/hal-00679842

G. Metegnier, T. Becking, M. A. Chebbi, G. I. Moumen, B. Schaack et al., Comparative paleovirological analysis of crustaceans identifies multiple widespread viral groups, Mobile DNA, vol.6, 2015.
URL : https://hal.archives-ouvertes.fr/hal-01204832

S. Lequime and L. Lambrechts, Discovery of flavivirus-derived endogenous viral elements in Anopheles mosquito genomes supports the existence of Anopheles-associated insect-specific flaviviruses, Virus Evol, vol.3, 2017.
URL : https://hal.archives-ouvertes.fr/hal-01445705

Y. Suzuki, L. Frangeul, L. B. Dickson, H. Blanc, Y. Verdier et al., Uncovering the repertoire of endogenous flaviviral elements in Aedes mosquito genomes, J Virol, vol.91, pp.571-588, 2017.
URL : https://hal.archives-ouvertes.fr/hal-01636504

A. Horst, J. C. Nigg, and B. W. Falk, Endogenous viral elements are widespread in arthropod genomes and commonly give rise to piRNAs, J Virol, 2018.

G. Antonelli and M. Pistello, Virology: A scientific discipline facing new challenges, Clin Microbiol Infect, vol.25, pp.133-135, 2019.

S. G. Black, F. Arnaud, M. Palmarini, and T. E. Spencer, Endogenous retroviruses in trophoblast differentiation and placental development, Am J Reprod Immunol, vol.64, p.20528833, 2010.
URL : https://hal.archives-ouvertes.fr/hal-01608531

J. Ashley, B. Cordy, D. Lucia, L. G. Fradkin, V. Budnik et al., Retrovirus-like Gag protein Arc1 binds RNA and traffics across synaptic boutons, Cell, vol.172, 2018.

E. D. Pastuzyn, C. E. Day, R. B. Kearns, M. Kyrke-smith, A. V. Taibi et al., The neuronal gene Arc encodes a repurposed retrotransposon Gag protein that mediates intercellular RNA transfer, Cell, vol.172, 2018.

B. J. Parker and J. A. Brisson, A laterally transferred viral gene modifies aphid wing plasticity, Curr Biol, vol.29, 2019.

M. R. Strand and G. R. Burke, Polydnaviruses: From discovery to current insights, Virology, 2015.

P. J. Krell and D. B. Stoltz, Unusual baculovirus of the parasitoid wasp Apanteles melanoscelus: Isolation and preliminary characterization, J Virol, vol.29, pp.1118-1130, 1979.

D. B. Stoltz, D. Guzo, E. R. Belland, C. J. Lucarotti, and E. A. Mackinnon, Venom promotes uncoating in vitro and persistence in vivo of DNA from a braconid polydnavirus, J Gen Virol, vol.69, pp.903-907, 1988.

A. Bezier, M. Annaheim, J. Herbiniere, C. Wetterwald, G. Gyapay et al., Polydnaviruses of braconid wasps derive from an ancestral nudivirus, Science, vol.323, pp.926-930, 2009.
URL : https://hal.archives-ouvertes.fr/hal-00402741

A. Volkoff, V. Jouan, S. Urbach, S. Samain, M. Bergoin et al., Analysis of virion structural components reveals vestiges of the ancestral ichnovirus genome, PLoS Pathog, vol.6, 2010.
URL : https://hal.archives-ouvertes.fr/inria-00537901

G. R. Burke, S. A. Thomas, J. H. Eum, and M. R. Strand, Mutualistic polydnaviruses share essential replication gene functions with pathogenic ancestors, PLoS Pathog, vol.9, 2013.

C. Bé-liveau, A. Cohen, D. Stewart, G. Periquet, A. Djoumad et al., Genomic and proteomic analyses indicate that Banchine and Campoplegine polydnaviruses have similar, if not identical, viral ancestors, J Virol, vol.89, pp.8909-8921, 2015.

W. N. Norton, S. B. Vinson, and D. B. Stoltz, Nuclear secretory particles associated with the calyx cells of the ichneumonid parasitoid Campoletis sonorensis (Cameron), Cell Tissue Res, vol.162, pp.195-208, 1975.

A. Volkoff, M. Ravallec, J. Bossy, P. Cerutti, J. Rocher et al., The replication of Hyposoter didymator polydnavirus: Cytopathology of the calyx cells in the parasitoid, Biol Cell, vol.83, pp.89926-89932, 1995.

A. Djoumad, D. Stoltz, C. Beliveau, B. Boyle, L. Kuhn et al., Ultrastructural and genomic characterization of a second banchine polydnavirus confirms the existence of shared features within this ichnovirus lineage, J Gen Virol, vol.94, p.23658210, 2013.

E. A. Herniou, E. Huguet, J. Theze, A. Bezier, G. Periquet et al., When parasitic wasps hijacked viruses: genomic and functional evolution of polydnaviruses, Philos Trans R Soc Lond B Biol Sci, vol.368, 2013.
URL : https://hal.archives-ouvertes.fr/hal-00864019

G. W. Blissard and G. F. Rohrmann, Baculovirus diversity and molecular biology, Ann Rev Entom, vol.35, pp.127-155, 1990.

A. Pichon, A. Bé-zier, S. Urbach, J. Aury, V. Jouan et al., Recurrent DNA virus domestication leading to different parasite virulence strategies, Sci Adv, vol.1, 2015.
URL : https://hal.archives-ouvertes.fr/hal-01318262

J. Young, E. Mackinnon, and P. Faulkner, The architecture of the virogenic stroma in isolated nuclei of Spodoptera frugiperda cells in vitro infected by Autographa californica nuclear polyhedrosis virus, J Struct Biol, vol.110, pp.141-153, 1993.

G. V. Williams and P. Faulkner, Cytological changes and viral morphogenesis during baculovirus infection. The Baculoviruses, pp.61-107, 1997.

G. W. Blissard and D. A. Theilmann, Baculovirus entry and egress from insect cells, Ann Rev Virol, vol.5, pp.113-139, 2018.

L. Deng and B. A. Webb, Cloning and expression of a gene encoding a Campoletis sonorensis polydnavirus structural protein, Arch Insect Biochem Physiol, vol.40, pp.30-40, 1999.

T. Nagamine, Y. Kawasaki, and S. Matsumoto, Induction of a subnuclear structure by the simultaneous expression of baculovirus proteins, IE1, LEF3, and P143 in the presence of hr, Virology, vol.352, pp.400-407, 2006.

Y. Shi, K. Li, P. Tang, Y. Li, Q. Zhou et al., Three-dimensional visualization of the Autographa californica multiple nucleopolyhedrovirus occlusion-derived virion envelopment process gives new clues as to its mechanism, Virology, vol.476, pp.298-303, 2015.

C. Suá-rez, S. Welsch, P. Chlanda, W. Hagen, S. Hoppe et al., Open membranes are the precursors for assembly of large DNA viruses, Cell Microbiol, 2013.

E. R. Quemin, S. Corroyer-dulmont, A. Baskaran, E. Penard, A. D. Gazi et al., Complex membrane remodeling during virion assembly of the 30,000-Year-Old mollivirus Sibericum, J Virol, vol.93, 2019.

D. B. Stoltz, C. Pavan, and A. Cunha, Nuclear polyhedrosis virus: A possible example of de novo intranuclear membrane morphogenesis, J Gen Virol, vol.19, pp.145-150, 1973.

S. Sun, V. B. Rao, and M. G. Rossmann, Genome packaging in viruses, Curr Opin Struct Biol, vol.20, pp.114-120, 2010.

G. F. Rohrmann, The AcMNPV genome: Gene content, conservation, and function, National Center for Biotechnology Information, vol.12, 2013.

M. Marek, O. Merten, L. Galibert, J. M. Vlak, and M. Oers, Baculovirus VP80 protein and the F-actin cytoskeleton interact and connect the viral replication factory with the nuclear periphery, J Virol, vol.85, pp.5350-5362, 2011.

T. Ohkawa and M. D. Welch, Baculovirus actin-based motility drives nuclear envelope disruption and nuclear egress, Curr Biol, vol.28, 2018.

L. Deng, D. B. Stoltz, and B. A. Webb, A gene encoding a polydnavirus structural polypeptide is not encapsidated, Virology, vol.269, pp.440-450, 2000.

A. Oomens and G. Blissard, Requirement for GP64 to drive efficient budding of Autographa californica multicapsid nucleopolyhedrovirus, Virology, vol.254, pp.297-314, 1999.

J. Zhou and G. W. Blissard, Identification of a GP64 subdomain involved in receptor binding by budded virions of the baculovirus Autographica californica multicapsid nucleopolyhedrovirus, J Virol, vol.82, pp.4449-4460, 2008.

V. Visconti, M. Eychenne, and I. Darboux, Modulation of antiviral immunity by the ichnovirus HdIV in Spodoptera frugiperda, Mol Immunol, vol.108, pp.89-101, 2019.
URL : https://hal.archives-ouvertes.fr/hal-02124769

A. Dobin, C. A. Davis, F. Schlesinger, J. Drenkow, C. Zaleski et al., STAR: ultrafast universal RNAseq aligner, Bioinformatics, vol.29, pp.15-21, 2012.

Y. Liao, G. K. Smyth, and W. Shi, featureCounts: an efficient general purpose program for assigning sequence reads to genomic features, Bioinformatics, vol.30, pp.923-930, 2013.

M. D. Robinson, D. J. Mccarthy, and G. K. Smyth, edgeR: a Bioconductor package for differential expression analysis of digital gene expression data, Bioinformatics, vol.26, pp.139-140, 2009.

S. Robin, M. Ravallec, M. Frayssinet, J. Whitfield, V. Jouan et al., Evidence for an ichnovirus machinery in parasitoids of coleopteran larvae, Virus Res, vol.263, pp.189-206, 2019.
URL : https://hal.archives-ouvertes.fr/hal-02059774

W. S. Rasband and . Imagej, National Institutes of Health, 1997.

T. Doré-mus, F. Cousserans, G. Gyapay, V. Jouan, P. Milano et al., Extensive transcription analysis of the Hyposoter didymator ichnovirus genome in permissive and non-permissive lepidopteran host species, PLoS ONE, vol.9, 2014.

G. Tellmann, The E-Method: a highly accurate technique for gene-expression analysis, Nat Methods, vol.3, 2006.

, R: A language and environment for statistical computing. R Foundation for Statistical Computing