K. Palavioux and F. Zenk, Noureddine El Mjiyad, and Damien Gironde (UMR1347, Dijon) for their technical support in the greenhouse experiments

N. Carlier, L. D. Corte, H. De-larambergue, H. Houtin, A. Klein et al., UMR1347, Dijon) for their help in plant phenotyping, Marie-Laure Pilet-Nayel (UMR IGEPP, Rennes), and Dominique Millot

, Dijon) for its help concerning the statistical analyses in R. This work was supported by the French ANR GENOPEA and the "INRA-BAP projet scientifique" SYMBIOPEA programs. The authors thank

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