B. H. Vanthoor, C. Stanghellini, E. J. Van-henten, and P. H. Visser, A methodology for model-based greenhouse design: Part 1, a greenhouse climate model for a broad range of designs and climates, Biosyst. Eng, vol.110, pp.363-377, 2011.

T. Boulard, J. Roy, J. Pouillard, H. Fatnassi, and A. Grisey, Modelling of micrometeorology, canopy transpiration and photosynthesis in a closed greenhouse using computational fluid dynamics, vol.158, pp.110-133, 2017.
URL : https://hal.archives-ouvertes.fr/hal-02392554

V. Truffault, S. Le-quillec, and E. Brajeul, Insights into the potential of semi-closed greenhouses and future perspectives for tomato crops, Acta Horti, vol.1227, pp.141-150, 2018.

, Concerning the Protection of Waters against Pollution Caused by Nitrates from Agricultural Sources, Council Directive 91/676/EEC of 12 December, p.20, 1991.

D. Savvas and N. Gruda, Application of soilless culture technologies in the modern greenhouse industry-A review, Eur. J. Hortic. Sci, vol.83, pp.280-293, 2018.

D. Schwarz, A. J. Thompson, and H. Kläring, Guidelines to use tomato in experiments with a controlled environment, Front. Plant Sci, vol.5, pp.625-641, 2014.

D. R. Hoagland, Inorganic nutrition of plants, Chronica Botanica Co, 1944.

C. De-kreij and W. Voogt, Van den Bos, A.; Baas, R. Voedingsoplossingen voor de teelt van tomaat in gesloten teeltsystemen, Brochure VG, 1997.

C. Bénard, H. Gautier, F. Bourgaud, D. Grasselly, B. Navez et al., Effects of low nitrogen supply on tomato (Solanum lycopersicum) fruit yield and quality with special emphasis on sugars, acids, ascorbate, carotenoids, and phenolic compounds, J. Agric. Food Chem, vol.57, pp.4112-4123, 2009.

G. Turcotte, R. Larouche, A. Carrier, and L. Lambert, Production de la tomate de serre au Québec (guide technique, p.15, 2017.

L. Bot, J. Adamowicz, S. Robin, P. Andriolo, J. L. Gary et al., Modelling nitrate uptake by greenhouse tomato crops at the short and long time scales. Acta Horti, vol.456, pp.237-246, 1998.

S. Adamowicz and J. Bot, Trends in modelling nitrate uptake. Acta Horti, vol.507, pp.231-240, 1991.

C. Sonneveld and W. Voogt, Nutrient Solutions for Soilless Cultures. In Plant Nutrition of Greenhouse Crops

. Springer, , pp.257-275, 2009.

L. Incrocci, D. Massa, and A. Pardossi, New Trends in the Fertigation Management of Irrigated Vegetable Crops, vol.3, p.37, 2017.

N. Bertin and M. Génard, Tomato quality as influenced by preharvest factors, Sci. Hortic, vol.233, pp.264-276, 2018.
URL : https://hal.archives-ouvertes.fr/hal-02629176

J. N. Davies, G. E. Hobson, and W. B. Mcglasson, The constituents of tomato fruit-the influence of environment, nutrition, and genotype, CRC Crit. Rev. Food Sci. Nutr, vol.15, pp.205-280, 1981.

E. A. Baldwin, J. W. Scott, M. A. Einstein, T. M. Malundo, B. T. Carr et al., Relationship between Sensory and Instrumental Analysis for Tomato Flavor, J. Amer. Soc. Hort. Sci, vol.123, pp.906-915, 1998.

D. Bhowmik, K. S. Kumar, S. Paswan, and S. Srivastava, Tomato-a natural medicine and its health benefits, J. Pharmacogn. Phytochem, vol.1, pp.33-43, 2012.

H. J. Klee and D. M. Tieman, The genetics of fruit flavour preferences, Nat. Rev. Genet, vol.19, pp.347-356, 2018.

H. Gautier, V. Diakou-verdin, C. Bénard, M. Reich, M. Buret et al., How Does Tomato Quality (Sugar, Acid, and Nutritional Quality) Vary with Ripening Stage, Temperature, and Irradiance?, J. Agr. Food Chem, vol.56, pp.1241-1250, 2008.

J. Ripoll, L. Urban, M. Staudt, F. Lopez-lauri, L. P. Bidel et al., Water shortage and quality of fleshy fruits-making the most of the unavoidable, J. Exp. Bot, vol.65, pp.4097-4117, 2014.
URL : https://hal.archives-ouvertes.fr/hal-01268584

J. Ripoll, L. Urban, and N. Bertin, The potential of the magic tom parental accessions to explore the genetic variability in tomato acclimation to repeated cycles of water deficit and recovery. Front, Plant Sci, 1187.
URL : https://hal.archives-ouvertes.fr/hal-01273701

Z. Wang, S. Li, and S. Malhi, Effects of fertilization and other agronomic measures on nutritional quality of crops, J. Sci. Food Agr, vol.88, pp.7-23, 2007.

C. Wang, F. Gu, J. Chen, H. Yang, J. Jiang et al., Assessing the response of yield and comprehensive fruit quality of tomato grown in greenhouse to deficit irrigation and nitrogen application strategies, Agr. Water Manage, vol.161, pp.9-19, 2015.

X. Wang and Y. Xing, Evaluation of the effects of irrigation and fertilization on tomato fruit yield and quality: A principal component analysis, vol.7, p.350, 2017.

N. Gruda, D. Savvas, G. Colla, and Y. Rouphael, Impacts of genetic material and current technologies on product quality of selected greenhouse vegetables -A review, Eur. J. Horticul. Sci, vol.83, pp.319-328, 2018.

R. Larbat, K. M. Olsen, R. Slimestad, T. Løvdal, C. Bénard et al., Influence of repeated short-term nitrogen limitations on leaf phenolics metabolism in tomato, Phytochemistry, vol.77, pp.119-128, 2012.

M. Parisi, L. Giordano, A. Pentangelo, B. D'onofrio, and G. Villari, Effects of different levels of nitrogen fertilization on yield and fruit quality in processing tomato, Acta Horticulturae, vol.700, pp.129-132, 2006.

T. Q. Zhang, C. S. Tan, K. Liu, C. F. Drury, A. P. Papadopoulos et al., Yield and Economic assessments of fertilizer nitrogen and phosphorus for processing tomato with drip fertigation, Agron. J, vol.102, pp.774-780, 2010.

F. Albornoz, Crop responses to nitrogen overfertilization: A review, Scientia Horticulturae, vol.205, pp.79-83, 2016.

V. Sarlikioti, P. H. Visser, and L. F. Marcelis, Exploring the spatial distribution of light interception and photosynthesis of canopies by means of a functional-structural plant model, Ann. Bot, vol.107, pp.875-883, 2011.

J. H. Kim, J. W. Lee, T. I. Ahn, J. H. Shin, K. S. Park et al., Sweet Pepper (Capsicum annuum L.) Canopy photosynthesis modeling using 3D plant architecture and light ray-tracing. Front, Plant Sci, vol.7, pp.1321-1331, 2016.

V. Sarlikioti, P. H. Visser, G. H. Buck-sorlin, and L. F. Marcelis, How plant architecture affects light absorption and photosynthesis in tomato: Towards an ideotype for plant architecture using a functional-structural plant model, Ann. Bot, vol.108, pp.1065-1073, 2011.
URL : https://hal.archives-ouvertes.fr/hal-01132291

C. Field, Allocating leaf nitrogen for the maximization of carbon gain: Leaf age as a control on the allocation program, Oecologia, vol.56, pp.341-347, 1989.

J. A. Prieto, G. Louarn, J. P. Peña, H. Ojeda, T. Simonneau et al., A leaf gas exchange model that accounts for intra-canopy variability by considering leaf nitrogen content and local acclimation to radiation in grapevine (Vitis vinifera L.), Plant Cell Environ, vol.35, pp.1313-1328, 2012.

K. Hikosaka, Optimality of nitrogen distribution among leaves in plant canopies, J. Plant Res, vol.129, pp.299-311, 2016.

J. R. Evans, Photosynthesis and nitrogen relationships in leaves of C3 plants, Oecologia, vol.78, pp.9-19, 1989.

M. Stitt, C. Müller, P. Matt, Y. Gibon, P. Carillo et al., Steps towards an integrated view of nitrogen metabolism, J. Exp. Bot, vol.53, pp.959-970, 2002.

L. Bot, J. Jeannequin, B. Fabre, and R. , Growth and Nitrogen Status of Soilless Tomato Plants Following Nitrate Withdrawal from the Nutrient Solution, Ann. Bot, vol.88, pp.361-370, 2001.

J. Vos and P. E. Van-der-putten, Effect of nitrogen supply on leaf growth, leaf nitrogen economy and photosynthetic capacity in potato, Field Crop. Res, vol.59, pp.63-72, 1998.

J. Vos, P. E. Putten, and C. J. Van-der-birch, Effect of nitrogen supply on leaf appearance, leaf growth, leaf nitrogen economy and photosynthetic capacity in maize, Zea mays L.). Field Crop. Res, vol.93, pp.64-73, 2005.

L. Guidi, G. Lorefice, A. Pardossi, F. Malorgio, F. Tognoni et al., Growth and photosynthesis of Lycopersicon esculentum (L.) plants as affected by nitrogen deficiency, Biol. Plantarum, vol.35, 1997.

T. Hirose and M. J. Werger, Maximizing daily canopy photosynthesis with respect to the leaf nitrogen allocation pattern in the canopy, Oecologia, vol.72, pp.520-526, 1987.

S. Hörtensteiner and U. Feller, Nitrogen metabolism and remobilization during senescence, J. Exp. Bot, vol.53, pp.927-937, 2002.

P. J. Hocking and B. T. Steer, The distribution and identity of assimilates in tomato with special reference to stem reserves, Ann. Bot, vol.73, pp.315-325, 1994.

W. V. Ende and D. Den-peshev, Sugars as antioxidants, Crop improvement under adverse conditions, pp.285-307, 2013.

W. Scheible, M. Lauerer, E. Schulze, M. Caboche, and M. Stitt, Accumulation of nitrate in the shoot acts as a signal to regulate shoot-root allocation in tobacco, Plant J, vol.11, pp.671-691, 1997.

C. Hermans, J. P. Hammond, P. J. White, and N. Verbruggen, How do plants respond to nutrient shortage by biomass allocation?, Trends Plant Sci, vol.11, pp.610-617, 2006.

M. J. Paul and S. P. Driscoll, Sugar repression of photosynthesis: The role of carbohydrates in signalling nitrogen deficiency through source:sink imbalance, Plant Cell Environ, vol.20, pp.110-116, 1997.

P. Muñoz, A. Antón, A. Paranjpe, J. Ariño, and J. I. Montero, High decrease in nitrate leaching by lower N input without reducing greenhouse tomato yield, Agron. Sustain. Dev, vol.28, pp.489-495, 2008.

L. Bot, J. Jeannequin, B. Fabre, and R. , Impacts of N-deprivation on the yield and nitrogen budget of rockwool grown tomatoes, Agronomie, vol.21, pp.341-350, 2001.
URL : https://hal.archives-ouvertes.fr/hal-00886122

C. Bénard, F. Bourgaud, and H. Gautier, Impact of temporary nitrogen deprivation on tomato leaf phenolics, Int. J. Mol. Sci, vol.12, pp.7971-7981, 2011.

E. Hoffland, M. J. Jeger, and M. L. Van-beusichem, Effect of nitrogen supply rate on disease resistance in tomato depends on the pathogen, Plant Soil, vol.218, pp.239-247, 2000.

L. Bot, J. Adamowicz, and S. , Nitrogen nutrition and use in horticultural crops, J. Crop Improv, vol.15, pp.323-367, 2006.

L. Wang, C. Qian, J. Bai, W. Luo, C. Jin et al., Difference in volatile composition between the pericarp tissue and inner tissue of tomato (Solanum lycopersicum) fruit. J. Food Process Pres, vol.42, 2017.

F. Wang, A. G. Smith, and M. L. Brenner, Temporal and spatial expression pattern of sucrose synthase during tomato fruit development, Plant Physiol, vol.104, pp.535-540, 1994.

M. M. Brown, J. L. Hall, and L. C. Ho, Sugar uptake by protoplasts isolated from tomato fruit tissues during various stages of fruit growth, Physiol. Plantarum, vol.101, pp.533-539, 1997.

Y. Cheng, T. Wang, J. Chen, and T. Lin, Spatial-temporal analyses of lycopene and sugar contents in tomatoes during ripening using chemical shift imaging, Postharvest Biology and Technology, vol.62, pp.17-25, 2011.

B. Van-de-poel, N. Vandenzavel, C. Smet, T. Nicolay, I. Bulens et al., Tissue specific analysis reveals a differential organization and regulation of both ethylene biosynthesis and E8 during climacteric ripening of tomato, BMC Plant Biol, vol.14, p.11, 2014.

B. Biais, C. Bénard, B. Beauvoit, S. Colombié, D. Prodhomme et al., Remarkable Reproducibility of Enzyme Activity Profiles in Tomato Fruits Grown under Contrasting Environments Provides a Roadmap for Studies of Fruit Metabolism, Plant Physiol, vol.164, pp.1204-1221, 2014.

S. Colombié, C. Nazaret, C. Bénard, B. Biais, V. Mengin et al., Modelling central metabolic fluxes by constraint-based optimization reveals metabolic reprogramming of developing Solanum lycopersicum (tomato) fruit, Plant J, vol.81, pp.24-39, 2015.

J. L. Rambla, Y. M. Tikunov, A. J. Monforte, A. G. Bovy, and A. Granell, The expanded tomato fruit volatile landscape, J Exp Bot, vol.65, pp.4613-4623, 2014.

L. Wang, E. A. Baldwin, and J. Bai, Recent Advance in aromatic volatile research in tomato fruit: The metabolisms and regulations, Food Bioproc. Technol, vol.9, pp.203-216, 2016.

S. Fishman and M. Génard, A biophysical model of fruit growth: Simulation of seasonal and diurnal dynamics of mass, Plant Cell Environ, vol.21, pp.739-752, 1998.

J. Fanwoua, P. H. Visser, E. De-heuvelink, X. Yin, P. C. Struik et al., A dynamic model of tomato fruit growth integrating cell division, cell growth and endoreduplication, Funct. Plant Biol, vol.40, pp.1098-1114, 2013.

B. Beauvoit, I. Belouah, N. Bertin, C. B. Cakpo, S. Colombié et al., Putting primary metabolism into perspective to obtain better fruits, Ann. Bot, vol.122, pp.1-21, 2018.
URL : https://hal.archives-ouvertes.fr/hal-02539159

A. Mozafar, Nitrogen fertilizers and the amount of vitamins in plants: A review, J. Plant Nutr, vol.16, pp.2479-2506, 1993.

H. Gautier, C. Massot, R. Stevens, S. Sérino, and M. Génard, Regulation of tomato fruit ascorbate content is more highly dependent on fruit irradiance than leaf irradiance, Ann. Bot, vol.103, pp.495-504, 2009.

E. Kaiser, A. Morales, J. Harbinson, J. Kromdijk, E. Heuvelink et al., Dynamic photosynthesis in different environmental conditions, J. Exp. Bot, vol.66, pp.2415-2426, 2015.

L. Gomez, E. Rubio, and M. Auge, A new procedure for extraction and measurement of soluble sugars in ligneous plants, J. Sci. Food Agr, vol.82, pp.360-369, 2002.

V. Truffault, G. Riqueau, C. Garchery, H. Gautier, and R. G. Stevens, Is monodehydroascorbate reductase activity in leaf tissue critical for the maintenance of yield in tomato?, J. Plant Physiol, vol.222, pp.1-8, 2018.
URL : https://hal.archives-ouvertes.fr/hal-01669403