D. L. Achat, L. Augusto, C. Morel, and M. R. Bakker, Predicting available phosphate ions from physical-chemical soil properties in acidic sandy soils under pine forests, J. Soils Sediments, vol.11, pp.452-466, 2011.
URL : https://hal.archives-ouvertes.fr/hal-02644025

D. L. Achat, M. R. Bakker, L. Augusto, E. Saur, L. Dousseron et al., Evaluation of the phosphorus status of P-deficient podzols in temperate pine stands: combining isotopic dilution and extraction methods, Biogeochemistry, vol.92, pp.183-200, 2009.
URL : https://hal.archives-ouvertes.fr/hal-02663717

D. L. Achat, C. Morel, M. R. Bakker, L. Augusto, S. Pellerin et al., Assessing turnover of microbial biomass phosphorus: combination of an isotopic dilution method with a mass balance model, Soil Biol. Biochem, vol.42, pp.2231-2240, 2010.
URL : https://hal.archives-ouvertes.fr/hal-02667318

R. Aerts, The freezer defrosting: global warming and litter decomposition rates in cold biomes, J.Ecol, vol.94, pp.713-724, 2006.

Q. Afnor and . Des-sols, Recueil de normes, vol.1, 1999.

K. Alef, P. Nannipieri, C. Trazar-cepeda, K. Alef, and P. Nannipieri, Phosphatase activity, Methods in Applied Soil Microbiology and Biochemistry, pp.338-339, 1995.

D. L. Achat, Phosphorus status of Siberian forest soils

K. Alef, K. Alef, and P. Nannipieri, Soil respiration, Methods in Applied Soil Microbiology and Biochemistry, pp.214-219, 1995.

F. Alt, Y. Oelmann, N. Herold, M. Schrumpf, and W. Wilcke, Phosphorus partitioning in grassland and forest soils of Germany as related to land-use type, management intensity, and land userelated pH, J. Plant Nutr. Soil Sci, vol.174, pp.195-209, 2011.

P. M. Attiwill and M. A. Adams, Nutrient cycling in forests, New Phytol, vol.124, pp.561-582, 1993.

L. Augusto, J. L. Dupouey, and J. Ranger, Effects of tree species on understory vegetation and environmental conditions in temperate forests, Ann. For. Sci, vol.60, pp.823-831, 2003.
URL : https://hal.archives-ouvertes.fr/hal-00883757

L. Augusto, J. Ranger, D. Binkley, R. , and A. , Impact of several common tree species of European temperate forests on soil fertility, Ann. For. Sci, vol.59, pp.233-253, 2002.
URL : https://hal.archives-ouvertes.fr/hal-02677150

B. Berg, M. P. Davey, A. De-marco, B. Emmett, M. Faituri et al., Factors influencing limit values for pine needle litter decomposition: a synthesis for boreal and temperate pine forest systems, Biogeochemistry, vol.100, pp.57-73, 2010.

J. A. Blanco, J. B. Imbert, and F. J. Castillo, Thinning affects Pinus sylvestris needle decomposition rates and chemistry differently depending on site conditions, Biogeochemistry, vol.106, pp.397-414, 2011.

P. C. Brookes, A. Landman, G. Pruden, and D. S. Jenkinson, Chloroform fumigation and the release of soil nitrogen: a rapid direct extraction method to measure microbial biomass nitrogen in soil, Soil Biol. Biochem, vol.17, pp.837-842, 1985.

E. K. Bünemann, A. Oberson, F. Liebisch, F. Keller, K. E. Annaheim et al., Rapid microbial phosphorus immobilization dominates gross phosphorus fluxes in a grassland soil with low inorganic phosphorus availability, Soil Biol. & Biochem, vol.51, pp.84-95, 2012.

L. Celi, E. Barberis, B. L. Turner, E. Frossard, and D. S. Baldwin, Abiotic stabilization of organic phosphorus in the environment, in: Organic Phosphorus in the Environment, pp.113-132, 2005.

C. R. Chen, S. Sinaj, L. M. Condron, E. Frossard, R. R. Sherlock et al., Characterization of phosphorus availability in selected New Zealand grassland soils, Nut. Cycl. Agroecosyst, vol.65, pp.89-100, 2003.

J. Chlachula, I. A. Dar, M. A. Dar, and . Intech-europe, Climate History and Early Peopling of Siberia, pp.495-538, 2011.

M. Chytry, J. Danihelka, S. Kubesova, P. Lustyk, N. Ermakov et al., Diversity of forest vegetation across a strong gradient of climatic continentality, Plant Ecol, vol.196, pp.61-83, 2008.

W. K. Cornwell, J. H. Cornelissen, K. Amatangelo, E. Dorrepaal, V. T. Eviner et al.,

S. D?az, E. Garnier, D. E. Gurvich, E. Kazakou, J. A. Klein et al., Plant species traits are the predominant control on litter decomposition rates within biomes worldwide, Ecol. Lett, vol.11, pp.1065-1071, 2008.

A. F. Cross and W. H. Schlesinger, A literature review and evaluation of the Hedley fractionation: applications to the biogeochemical cycle of soil phosphorus in natural ecosystems, Geoderma, vol.64, pp.197-214, 1995.

M. A. De-graaff, K. J. Van-groenigen, J. Six, B. Hungate, and C. Van-kessel, Interactions between plant growth and soil nutrient cycling under elevated CO 2 : a meta-analysis, Glob. Change Biol, vol.12, pp.2077-2091, 2006.

A. De-schrijver, L. Vesterdal, K. Hansen, P. De-frenne, L. Augusto et al., Four decades of post-agricultural forest development have caused major redistributions of soil phosphorus fractions, Oecologia, vol.169, pp.221-234, 2012.
URL : https://hal.archives-ouvertes.fr/hal-02650056

F. Dijkstra and W. Cheng, Increased soil moisture content increases plant N uptake and the abundance of 15 N in plant biomass, Plant Soil, vol.302, pp.263-271, 2008.

J. J. Elser, M. E. Bracken, E. E. Cleland, D. S. Gruner, W. S. Harpole et al., Global analysis of nitrogen and phosphorus limitation of primary producers in freshwater, marine and terrestrial ecosystems, Ecol. Lett, vol.10, pp.1135-1142, 2007.

J. J. Elser, A world awash with nitrogen, Science, vol.334, pp.1504-1505, 2011.

J. C. Fardeau, Dynamics of phosphate in soils. An isotopic outlook, Fert. Res, vol.45, pp.91-100, 1996.

J. C. Fardeau, Le phosphore assimilable des sols: sa représentation par un modèle fonctionnelà plusieurs compartiments, Agronomie, vol.13, pp.317-331, 1993.

R. F. Fisher and D. Binkley, Food and Agriculture Organization: Global Forest Resources Assessment 2005, Progress towards sustainable forest management, FAO Forestry paper 147, vol.489, 2000.

E. Frossard, S. ;. Sinaj, P. , and Z. , The isotope exchange kinetic technique: a method to describe the availability of inorganic nutrients, Applications to, vol.33, pp.61-77, 1997.

E. Frossard, J. W. Stewart, and R. J. St-arnaud, Distribution and mobility of phosphorus in grassland and forest soils of Saskatchewan, Can. J. Soil Sci, vol.69, pp.401-416, 1989.

R. Giesler, T. Petersson, and P. Högberg, Phosphorus Limitation in Boreal Forests: Effects of Aluminum and Iron Accumulation in the Humus Layer, Ecosystems, vol.5, pp.300-314, 2002.

R. Giesler, F. Satoh, U. Ilstedt, and A. Nordgren, Microbially available phosphorus in boreal forests: effects of aluminium and iron accumulation in the humus layer, Ecosystems, vol.7, pp.208-217, 2004.

N. Gilbert, The disappearing nutrient, Nature, vol.461, pp.716-718, 2009.

P. M. Groffman, C. T. Driscoll, T. J. Fahey, J. P. Hardy, R. D. Fitzhugh et al., Colder soils in a warmer world: A snow manipulation study in a northern hardwood forest ecosystem, Biogeochemistry, vol.56, pp.135-150, 2001.

D. L. Achat, Phosphorus status of Siberian forest soils, p.751

A. Guckland, M. Jacob, H. Flessa, F. M. Thomas, and C. Leuschner, Acidity, nutrient stocks, and organic-matter content in soils of a temperate deciduous forest with different abundance of European beech (Fagus sylvatica L.), J. Plant Nutr. Soil Sc, vol.172, pp.500-511, 2009.

K. Hansen, L. Vesterdal, I. K. Schmidt, P. Gundersen, L. Sevel et al., Litterfall and nutrient return in five tree species in a common garden experiment, Forest Ecol. Manag, vol.257, pp.2133-2144, 2009.

W. S. Harpole, J. T. Ngai, E. E. Cleland, E. W. Seabloom, E. T. Borer et al., Ecol Lett, vol.14, pp.852-862, 2011.

I. P. Hartley, D. W. Hopkins, M. Sommerkorn, and P. A. Wookey, The response of organic matter mineralisation to nutrient and substrate additions in sub-arctic soils, Soil Biol. Biochem, vol.42, pp.92-100, 2010.

S. Hättenschwiler and P. Gasser, Soil animals alter plant litter diversity effects on decomposition, vol.102, pp.1519-1524, 2005.

L. O. Hedin, E. N. Jack-brookshire, D. N. Menge, and A. R. Barron, The Nitrogen Paradox in Tropical Forest Ecosystems, Annu. Rev. Ecol. Evol. S, vol.40, pp.613-648, 2009.

P. Hietz, B. L. Turner, W. Wanek, A. Richter, C. A. Nock et al., Long-term change in the nitrogen cycle of tropical forests, Science, vol.334, pp.664-666, 2011.

P. Hinsinger, Bioavailability of soil inorganic P in the rhizosphere as affected by root-induced chemical changes: a review, Plant Soil, vol.237, pp.173-195, 2001.
URL : https://hal.archives-ouvertes.fr/hal-02679491

G. W. Holtgrieve, D. E. Schindler, W. O. Hobbs, P. R. Leavitt, E. J. Ward et al., A coherent signature of anthropogenic nitrogen deposition to remote watersheds of the northern hemisphere, Science, vol.334, pp.1545-1548, 2011.

M. A. Huston and S. Wolverton, The global distribution of net primary production: resolving the paradox, Ecol. Monogr, vol.79, pp.343-377, 2009.

I. P. Ipcc, Fourth Assessment Report: Climate Change, 2007.

C. M. Iversen, Digging deeper: fine-root responses to rising atmospheric CO 2 concentration in forested ecosystems, New Phytol, vol.186, pp.346-357, 2010.

R. B. Jackson, J. Canadell, J. R. Ehleringer, H. A. Mooney, O. A. Sala et al., A global analysis of root distributions for terrestrial biomes, Oecologia, vol.108, pp.389-411, 1996.

A. H. Johnson, J. Frizano, D. R. Vann, A. Jungk, and N. Claassen, Biogeochemical implications of labile phosphorus in forest soils determined by the Hedley fractionation procedure, Adv. Agron, vol.135, pp.53-110, 1997.

H. Lambers, J. A. Raven, G. R. Shaver, and S. E. Smith, Plant nutrient-acquisition strategies change with soil age, Trends Ecol. Evol, vol.23, pp.95-103, 2008.

N. N. Lashchinskiy, Vegetation of Salair mountain range, 2009.

H. Li, Y. Han, and Z. Cai, Nitrogen mineralization in paddy soils of the Taihu Region of China under anaerobic conditions: dynamics and model fitting, Geoderma, vol.115, pp.161-175, 2003.

D. Loustau, A. Bosc, A. Colin, J. Ogée, H. Davi et al., Modeling climate change effects on the potential production of French plains forests at the subregional level, Tree Physiol, vol.25, pp.813-823, 2005.
URL : https://hal.archives-ouvertes.fr/hal-02682718

Y. Luo, D. Hui, and D. Zhang, Elevated CO 2 stimulates net accumulations of carbon and nitrogen in land ecosystems: a metaanalysis, Ecology, vol.87, pp.53-63, 2006.

J. A. Mckeague and J. H. Day, Dithionite and oxalate-extractable Fe and Al as aids in differentiating various classes of soils, Can. J. Soil Sci, vol.46, pp.13-22, 1966.

C. J. Mikan, J. P. Schimel, A. P. Doyle, R. Hassan, R. Scholes et al., Temperature controls of microbial respiration in arctic tundra soils above and below freezing, Soil Biol. Biochem, vol.34, p.838, 2002.

C. Morel and C. Plenchette, Is the isotopically echangeable phosphate of a loamy soil the plant-available P?, Plant Soil, vol.158, pp.287-297, 1994.

C. Morel, H. Tiessen, and J. W. Stewart, Correction for Psorption in the measurement of soil microbial biomass P by CHCl 3 fumigation, Soil Biol. Biochem, vol.28, pp.1699-1706, 1996.
URL : https://hal.archives-ouvertes.fr/hal-02699116

C. Morel, H. Tunney, D. Plénet, and S. Pellerin, Transfer of Phosphate Ions between Soil and Solution: Perspectives in Soil Testing, J. Environ. Qual, vol.29, pp.20-59, 2000.
URL : https://hal.archives-ouvertes.fr/hal-02699114

C. Morel, Mobilité et biodisponibilité du phosphore dans les sols cultivés: mécanismes, modélisation et diagnostic, Oceanis, vol.33, pp.51-74, 2007.

W. Negassa and P. Leinweber, How does the Hedley sequential phosphorus fractionation reflect impacts of land use and management on soil phosphorus: A review, J. Plant Nutr. Soil Sci, vol.172, pp.305-325, 2009.

S. Nobrega and P. Grogan, Deeper Snow Enhances Winter Respiration from Both Plant-associated and Bulk Soil Carbon Pools in Birch Hummock Tundra, Ecosystems, vol.10, pp.419-431, 2007.

R. J. Norby, E. H. Delucia, B. Gielen, C. Calfapietra, C. P. Giardina et al., , vol.102, pp.18052-18056, 2005.

R. Oren, D. S. Ellsworth, K. H. Johnsen, N. Phillips, B. E. Ewers et al., Soil fertility limits carbon sequestration by forest ecosystems in a CO 2 -enriched atmosphere, Nature, vol.411, pp.469-472, 2001.

D. J. Read, J. R. Leake, and J. Perez-moreno, Mycorrhizal fungi as drivers of ecosystem processes in heathland and boreal forest biomes, Can. J. Bot, vol.82, pp.1243-1263, 2004.

P. B. Reich, S. E. Hobbie, T. Lee, D. S. Ellsworth, J. B. West et al., Nitrogen limitation constrains sustainability of ecosystem response to CO 2, Nature, vol.440, pp.922-925, 2006.

A. E. Richardson and R. J. Simpson, Soil Microorganisms Mediating Phosphorus Availability, Plant Physiol, vol.156, pp.989-996, 2011.

D. L. Achat, Phosphorus status of Siberian forest soils

W. M. Saunders and E. G. Williams, Observations on the determination of total organic phosphorus in soils, J. Soil Sci, vol.6, pp.247-267, 1955.

J. K. Shuman, H. H. Shugart, and T. Halloran, L: Sensitivity of Siberian larch forests to climate change, Global Change Biol, vol.17, pp.2370-2384, 2011.

R. Strauss, G. W. Brümmer, and N. J. Barrow, Effects of crystallinity of goethite: II. rates of sorption and desorption of phosphate, Terrestrial biozones: available online, vol.48, pp.101-114, 1997.

S. M. Thomas, D. Whitehead, J. B. Reid, F. J. Cook, J. A. Adams et al., Growth, loss, and vertical distribution of Pinus radiata fine roots growing at ambient and elevated CO 2 concentration, Global Change Biol, vol.5, pp.107-121, 1999.

P. Trichet, M. R. Bakker, L. Augusto, P. Alazard, and D. Merzeau, Fifty years of pine fertilization experiments in the Landes of Gascogne (France), Forest Sci, vol.55, pp.390-402, 2009.

B. L. Turner, R. Baxter, N. Mahieu, S. Sjögersten, and B. A. Whitton, Phosphorus compounds in subarctic Fennoscandian soils at the mountain birch (Betula pubescens)-tundra ecotone, Soil Biol. Biochem, vol.36, pp.815-823, 2004.

B. L. Turner, B. J. Cade-menun, L. M. Condron, and S. Newman, Extraction of soil organic phosphorus, Talanta, vol.66, pp.294-306, 2005.

M. Ushio, K. Kitayama, T. C. Balser, P. P. Van-veldhoven, and G. P. Mannaerts, Tree species effects on soil enzyme activities through effects on soil physicochemical and microbial properties in a tropical montane forest on Mt, Anal. Biochem, vol.53, pp.45-48, 1987.

E. D. Vance, P. C. Brookes, and D. S. Jenkinson, An extraction method for measuring soil microbial biomass C, Soil Biol. Biochem, vol.19, pp.703-707, 1987.

G. Vazquez, G. Antorrena, J. Gonzalez, and S. Freire, Studies on the composition of Pinus Pinaster foliage, Bioresource Technol, vol.51, pp.83-87, 1995.

A. Vincent, J. Schleucher, G. Gröbner, J. Vestergren, P. Persson et al., Changes in organic phosphorus composition in boreal forest humus soils: the role of iron and aluminium, vol.108, pp.485-499, 2012.

P. M. Vitousek, S. Porder, B. Z. Houlton, and O. A. Chadwick, Terrestrial phosphorus limitation: mechanisms, implications, and nitrogen-phosphorus interactions, Ecol. Appl, vol.20, pp.5-15, 2010.

J. Voriskova, P. Dobiasova, J. Snajdr, D. Vanek, T. Cajthaml et al., Chemical composition of litter affects the growth and enzyme production by the saprotrophic basidiomycete Hypholoma fasciculare, Fungal Ecol, vol.4, pp.417-426, 2011.

M. R. Walbridge, C. J. Richardson, and W. T. Swank, Vertical distribution of biological and geochemical phosphorus subcycles in two Appalachian forest soils, Biogeochemistry, vol.13, pp.61-85, 1991.

Y. P. Wang, B. Z. Houlton, and C. B. Field, A model of biogeochemical cycles of carbon, nitrogen, and phosphorus including symbiotic nitrogen fixation and phosphatase production, Global Biogeochem. Cy, vol.21, 1018.