C. Gebhardt, The historical role of species from the Solanaceae plant family in genetic research, Theor. Appl. Genet, vol.129, pp.2281-2294, 2016.

R. G. Olmstead, L. Bohs, H. A. Migid, E. Santiago-valentin, V. F. Garcia et al., A molecular phylogeny of the Solanaceae, Taxon, vol.57, pp.1159-1181, 2008.

B. Singh, Survey and indexing of weeds growing around potato fields for their role as an inoculum source for Potato leafroll virus (PLRV), Br. Biotechnol. J, vol.16, pp.1-8, 2016.

M. W. Beijerinck, Concerning a contagium vivum fluidum as cause of the spot disease of tobacco leaves, Phytopatologial Classics

J. Johnson, , vol.7, pp.33-52, 1942.

K. B. Scholthof, Tobacco mosaic virus: A model system for plant biology, Annu. Rev. Phytopathol, vol.42, pp.13-34, 2004.

, ICTV Master Species List 2018b.v2. Available online, 2019.

A. Pecman, D. Kutnjak, I. Gutiérrez-aguirre, I. Adams, A. Fox et al., Next generation sequencing for detection and discovery of plant viruses and viroids: Comparison of two approaches, Front. Microbiol, vol.8, 1998.

D. E. Villamor, T. Ho, M. Al-rwahnih, R. R. Martin, and I. E. Tzanetakis, High throughput sequencing for plant virus detection and discovery, Phytopathology, vol.109, pp.716-725, 2019.

C. Xu, X. Sun, A. Taylor, C. Jiao, Y. Xu et al., Diversity, distribution, and evolution of tomato viruses in China uncovered by small RNA sequencing, J. Virol, vol.91, pp.173-190, 2017.

L. Lotos, A. Olmos, C. Orfanidou, K. Efthimiou, A. Avgelis et al., Insights into the etiology of polerovirus-induced pepper yellows disease, Phytopathology, vol.107, pp.1567-1576, 2017.

J. C. Ng and K. L. Perry, Transmission of plant viruses by aphid vectors, Mol. Plant Pathol, vol.5, pp.505-511, 2004.

S. Mondal, E. J. Wenninger, P. J. Hutchinson, J. L. Whitworth, D. Shrestha et al., Responses of aphid vectors of Potato leaf roll virus to potato varieties, Plant Dis, vol.101, pp.1812-1818, 2017.

D. Rotenberg, A. L. Jacobson, D. J. Schneweis, and A. E. Whitfield, Thrips transmission of tospoviruses, Curr. Opin. Virol, vol.15, pp.80-89, 2015.

H. Pospieszny, M. Budziszewska, B. Hasiów-jaroszewska, A. Obr?palska-st?plowska, and N. Borodynko, Biological and molecular characterization of Polish isolates of Tomato torrado virus, J. Phytopathol, vol.158, pp.56-62, 2010.

M. Verbeek, P. J. Van-bekkum, A. M. Dullemans, and R. A. Van-der-vlugt, Torradoviruses are transmitted in a semi-persistent and stylet-borne manner by three whitefly vectors, Virus Res, vol.186, pp.55-60, 2014.

E. Riga, R. Larsen, K. Eastwell, N. Guerra, L. Guerra et al., Rapid detection of Tobacco rattle tobravirus in viruliferous Paratrichodorus allius from greenhouse and field specimens, J. Nematol, vol.41, pp.60-63, 2009.

M. L. Adams, Transmission of plant viruses by fungi, Ann. Appl. Biol, vol.118, pp.479-492, 1991.

B. Kassanis and I. Macfarlane, Transmission of tobacco necrosis virus by zoospores of Olpidium brassicae, J. Gen. Microbiol, vol.36, pp.79-93, 1964.

R. A. Jones and B. D. Harrison, The behaviour of potato mop-top virus in soil, and evidence for its transmission by Spongospora subterranea (Wallr.) Lagerh, Ann. Appl. Biol, vol.63, pp.1-17, 1969.

A. Alfaro-fernández, M. Del-carmen-córdoba-sellés, J. Herrera-vásquez, M. Cebrián, and C. Jordá, Transmission of Pepino mosaic virus by the fungal vector Olpidium virulentus, J. Phytopathol, vol.158, pp.217-226, 2010.

E. Deja-sikora, L. Mercy, C. Baum, and K. Hrynkiewicz, The contribution of endomycorrhiza to the performance of Potato virus Y-infected solanaceous plants: Disease alleviation or exacerbation?, Front. Microbiol, vol.10, 2019.

A. Dombrovsky, E. Smith, J. C. Jimenez-lopez, and . Ed, Seed transmission of tobamoviruses: Aspects of global disease distribution, Advances in Seed Biology, pp.234-260, 2017.

I. M. Hanssen, R. Mumford, D. Blystad, I. Cortez, B. Hasiów-jaroszewska et al., Seed transmission of Pepino mosaic virus in tomato, Eur. J. Plant Pathol, vol.126, pp.145-152, 2010.

H. Pospieszny, N. Borodynko-filas, B. Hasiów-jaroszewska, N. Rymelska, and S. F. Elena, Transmission rate of two Polish Tomato torrado virus isolates through tomato seeds, J. Gen. Plant Pathol, vol.85, pp.109-115, 2019.

M. A. Aranda and J. Freitas-astua, Ecology and diversity of plant viruses, and epidemiology of plant virus-induced diseases, Ann. Appl. Biol, vol.171, pp.1-4, 2017.

P. K. Anderson, A. A. Cunningham, N. G. Patel, F. J. Morales, P. R. Epstein et al., Emerging infectious diseases of plants: Pathogen pollution, climate change and agrotechnology drivers, Trends Ecol. Evol, vol.19, pp.535-544, 2004.

I. M. Hanssen, M. Lapidot, and B. P. Thomma, Emerging viral diseases of tomato crop, Mol. Plant Microbe Interact, vol.23, pp.539-548, 2010.

N. Salem, A. Mansour, M. Ciuffo, B. W. Falk, and M. Turina, A new tobamovirus infecting tomato crops in Jordan, Arch. Virol, vol.161, pp.503-506, 2016.

M. R. Rojas, R. L. Gilbertson, and M. J. Roossinck, Emerging plant viruses: A diversity of mechanisms and opportunities, Plant Virus Evolution, pp.27-51, 2008.

N. Luria, E. Smith, V. Reingold, I. Bekelman, M. Lapidot et al., A new Israeli tobamovirus isolate infects tomato plants harboring Tm-2 2 resistance genes, PLoS ONE, vol.12, 2017.

K. Ling, T. Tian, S. Gurung, R. Salati, and A. Gilliard, First report of tomato brown rugose fruit virus infecting greenhouse tomato in the United States

K. Just, W. N. Leke, M. N. Sattar, A. Luik, and A. Kvarnheden, Detection of Tomato yellow leaf curl virus in imported tomato fruit in northern Europe, Plant Pathol, vol.63, pp.1454-1460, 2014.

R. A. Jones, Future scenarios for plant virus pathogens as climate change progresses, Adv. Virus Res, vol.95, pp.87-147, 2016.

R. A. Jones, Virus disease problems facing potato industries worldwide: Viruses found, climate change implications, rationalizing virus strain nomenclature and addressing the Potato virus Y issue, The Potato: Botany, Production and Uses, pp.202-224, 2014.

I. Cooper and R. A. Jones, Wild plants and viruses: Under-investigated ecosystems, Adv. Virus Res, vol.67, pp.1-47, 2006.

R. A. Jones and B. A. Coutts, Spread of introduced viruses to new plants in natural ecosystems and the threat this poses to plant biodiversity, Mol. Plant Pathol, vol.16, pp.541-545, 2015.

M. Forsius, S. Anttila, L. Arvola, I. Bergström, H. Hakola et al., Impacts and adaptation options of climate change on ecosystem services in Finland: A model based study, Curr. Opin. Environ. Sustain, vol.5, pp.26-40, 2013.

J. J. Burdon and P. H. Thrall, Pathogen evolution across the agro-ecological interface: Implications for disease management, Evol. Appl, vol.1, pp.57-65, 2008.

J. E. Duffus, Role of weeds in the incidence of virus diseases, Annu. Rev. Phytopathol, vol.9, pp.319-340, 1971.

I. D. Bedford, A. Kelly, G. K. Banks, R. W. Briddon, J. L. Cenis et al., Solanum nigrum: An indigenous weed reservoir for a tomato yellow leaf curl geminivirus in southern Spain, Eur. J. Plant Pathol, vol.104, pp.221-222, 1998.

M. J. Roossinck and F. García-arenal, Ecosystem simplification, biodiversity loss and plant virus emergence, Curr. Opin. Virol, vol.10, pp.56-62, 2015.

M. J. Roossinck, Plants, viruses and the environment: Ecology and mutualism, Virology, vol.479, pp.271-277, 2015.

S. H. Koh, H. Li, K. Sivasithamparam, R. Admiraal, M. G. Jones et al., Evolution of a wild-plant tobamovirus passaged through an exotic host: Fixation of mutations and increased replication

M. Lapidot, J. P. Legg, W. M. Wintermantel, and J. E. Polston, Management of whitefly-transmitted viruses in open-field production systems, Adv. Virus Res, vol.90, pp.147-206, 2014.

V. Nicaise, Crop immunity against viruses: Outcomes and future challenges. Front, Plant Sci, vol.5, 2014.
URL : https://hal.archives-ouvertes.fr/hal-02638251

B. Dupuis, J. Cadby, G. Goy, M. Tallant, J. Derron et al., Control of potato virus Y (PVY) in seed potatoes by oil spraying, straw mulching and intercropping, Plant Pathol, vol.66, pp.960-969, 2017.

J. M. Lanter, J. M. Mcguire, and M. J. Goode, Persistence of tomato mosaic virus in tomato debris and soil under field conditions, Plant Dis, vol.66, pp.552-555, 1982.

R. A. Jones and M. J. Barbetti, Influence of climate change on plant disease infections and epidemics caused by viruses and bacteria. CAB Rev, vol.7, pp.1-31, 2012.

A. N. Laws and A. Joern, Ecological mechanisms underlying arthropod species diversity in grasslands, Annu. Rev. Entomol, vol.58, pp.19-36, 2013.

T. M. Shates, P. Sun, C. Malmstrom, C. Dominguez, and K. Mauck, Addressing research needs in the field of plant virus ecology by defining knowledge gaps and developing wild dicot study systems, Front. Microbiol, vol.9, p.3305, 2019.

I. Pagán and E. C. Holmes, Long-term evolution of the Luteoviridae: Time scale and mode of virus speciation, J. Virol, vol.84, pp.6177-6187, 2010.

A. J. Gibbs, J. Wood, F. García-arenal, K. Ohshima, and J. S. Armstrong, Tobamoviruses have probably co-diverged with their eudicotyledonous hosts for at least 110 million years

A. J. Gibbs, K. Ohshima, M. J. Phillips, and M. J. Gibbs, The prehistory of potyviruses: Their initial radiation was during the dawn of agriculture, PLoS ONE, vol.3, 2008.

E. P. Rybicki, A Top Ten list for economically important plant viruses, Arch. Virol, vol.160, pp.17-20, 2015.

K. B. Scholthof, S. Adkins, H. Czosnek, P. Palukaitis, E. Jacquot et al., Top 10 plant viruses in molecular plant pathology, Mol. Plant. Pathol, vol.9, pp.938-954, 2011.
URL : https://hal.archives-ouvertes.fr/hal-02652176

M. Fuchs, Plant resistance to viruses: Engineered resistance, Encyclopedia of Virology, pp.156-164, 2008.

A. Fraile, J. L. Alonso-prados, M. A. Aranda, J. J. Bernal, J. M. Malpica et al., Genetic exchange by recombination or reassortment is infrequent in natural populations of a tripartite RNA plant virus, J. Virol, vol.71, pp.934-940, 1997.

S. García-andrés, F. Monci, J. Navas-castillo, and E. Moriones, Begomovirus genetic diversity in the native plant reservoir Solanum nigrum: Evidence for the presence of a new virus species of recombinant nature, Virology, vol.350, pp.433-442, 2006.

D. B. Paudel and H. Sanfaçon, Exploring the diversity of mechanisms associated with plant tolerance to virus infection. Front, Plant Sci, vol.9, 1575.

P. Xu, F. Chen, J. P. Mannas, T. Feldman, L. W. Sumner et al., Virus infection improves drought tolerance, New Phytol, vol.180, pp.911-921, 2008.

M. J. Roossinck, The good viruses: Viral mutualistic symbioses, Nat. Rev. Microbiol, vol.9, pp.99-108, 2011.

M. J. Roossinck, A new look at plant viruses and their potential beneficial roles in crops, Mol. Plant Pathol, vol.16, pp.331-333, 2015.

T. Fukuhara, M. Tabara, H. Koiwa, and H. Takahashi, Effect of asymptomatic infection with southern tomato virus on tomato plants, Arch. Virol, vol.2020, pp.11-20

K. Pechinger, K. M. Chooi, R. M. Macdiarmid, S. J. Harper, and H. Ziebell, A new era for mild strain cross-protection, Viruses, vol.11, 2019.

T. Matsumoto, Y. Nara, H. Furuya, H. Takahashi, K. Tairako et al., Characteristics for practical use of attenuated isolate UA-Fukushima of Tomato mosaic virus, J. Gen. Plant Pathol, vol.68, pp.382-384, 2002.

G. Chewachong, S. Miller, J. Blakeslee, D. Francis, T. Morris et al., Generation of an attenuated, cross-protective Pepino mosaic virus variant through alignment-guided mutagenesis of the viral capsid protein, Phytopathology, vol.105, pp.126-134, 2015.

I. M. Hanssen, I. Gutierrez-aguirre, A. Paeleman, K. Goen, L. Wittemans et al., Cross-protection or enhanced symptom display in greenhouse tomato co-infected with different Pepino mosaic virus isolates, Plant Pathol, vol.59, pp.13-21, 2010.

B. Hasiów-jaroszewska, J. Minicka, and H. Pospieszny, Cross-protection between different pathotypes of Pepino mosaic virus representing Chilean 2 genotype, Acta Sci. Pol. Hortorum Cultus, vol.13, pp.177-185, 2014.

J. Agüero, C. Gómez-aix, R. N. Sempere, J. García-villalba, J. García-núñez et al., Stable and broad spectrum cross-protection against Pepino mosaic virus attained by mixed infection. Front, Plant Sci, vol.9, 1810.

A. Grupa and J. Syller, Cross-protection between a naturally occurring mild isolate of Potato virus M (PVM) and a more virulent isolate in Datura metel plants, J. Phytopathol, vol.164, pp.69-73, 2016.

T. J. Hall, Resistance at the TM-2 locus in the tomato to tomato mosaic virus, Euphytica, vol.29, pp.189-197, 1980.

V. L. Calder and P. Palukaitis, Nucleotide sequence analysis of the movement genes of resistance breaking strains of tomato mosaic virus, J. Gen. Virol, vol.73, pp.165-168, 1992.

F. C. Lanfermeijer, J. Warmink, and J. Hille, The products of the broken Tm-2 and the durable Tm-22 resistance genes from tomato differ in four amino acids, J. Exp. Bot, vol.56, pp.2925-2933, 2005.

A. Niehl, M. Soininen, M. M. Poranen, and M. Heinlein, Synthetic biology approach for plant protection using dsRNA, Plant Biotechnol. J, vol.16, pp.1679-1687, 2018.
URL : https://hal.archives-ouvertes.fr/hal-02342530

A. M. Hamilton, On three new virus diseases of hyoscyamus niger, Ann. Appl. Biol, vol.19, pp.550-567, 1932.

O. Lovisolo and R. Bartels, On a new strain of henbane mosaic virus from Physalis alkekengi, J. Phytopathol, vol.69, pp.189-201, 1970.

A. Pecman, D. Kutnjak, N. Mehle, M. T. ?nidari?, I. Gutiérrez-aguirre et al., High-throughput sequencing facilitates characterization of a "forgotten" plant virus: The case of a Henbane mosaic virus infecting tomato, Front. Microbiol, vol.9, p.2739, 2018.

B. Moury, F. Fabre, E. Hébrard, and R. Froissart, Determinants of host species range in plant viruses, J. Gen. Virol, vol.98, pp.862-873, 2017.
URL : https://hal.archives-ouvertes.fr/hal-02739696

C. G. Orfanidou, P. G. Pappi, K. E. Efthimiou, N. I. Katis, and V. I. Maliogka, Transmission of Tomato chlorosis virus (ToCV) by Bemisia tabaci biotype Q and evaluation of four weed species as viral sources, Plant Dis, vol.100, pp.2043-2049, 2016.

M. J. Mcleisch, A. Fraile, and F. García-arenal, Evolution of plant-virus interactions: Host range and virus emergence, Curr. Opin. Virol, vol.34, pp.50-55, 2019.

K. S. Sastry, B. Mandal, J. Hammond, S. W. Scott, and R. W. Briddon, Encyclopedia of Plant Viruses and Viroids, 2019.

T. Ictv-virus, , 2019.

, VIDE Database: Plant Viruses Online. Available online, 2020.

. Ncbi-virus-portal, , 2020.

S. Bag, N. Mitter, S. Eid, and H. R. Pappu, Complementation between two tospoviruses facilitates the systemic movement of a plant virus silencing suppressor in an otherwise restrictive host, PLoS ONE, vol.7, 2012.

A. Hassani-mehraban, A. Brenkman, N. Broek, R. Goldbach, and R. Kormelink, RNAi-mediated transgenic tospovirus resistance broken by intraspecies silencing suppressor protein complementation, Mol. Plant Microbe Interact, vol.22, pp.1250-1257, 2009.

S. Sánchez-campos, G. Domínguez-huerta, L. Díaz-martínez, D. M. Tomás, J. Navas-castillo et al., Differential shape of geminivirus mutant spectra across cultivated and wild hosts with invariant viral consensus sequences. Front, Plant Sci, vol.9, p.932, 2018.

M. Juárez, M. P. Rabadán, L. D. Martínez, M. Tayahi, A. Grande-pérez et al., Natural hosts and genetic diversity of the emerging Tomato leaf curl New Delhi virus in Spain, Front. Microbiol, vol.10, 2019.

Y. Ma, A. Marais, M. Lefebvre, C. H. Faure, and T. Candresse, Metagenomic analysis of virome cross-talk between cultivated Solanum lycopersicum and wild Solanum nigrum, Virology, vol.540, pp.38-44, 2020.
URL : https://hal.archives-ouvertes.fr/hal-02625134

S. A. Bratsch, S. Grinstead, T. C. Creswell, G. E. Ruhl, and D. Mollov, Characterization of Tomato necrotic spot virus, a subgroup 1 ilarvirus causing necrotic foliar, stem, and fruit symptoms in tomatoes in the United States, Plant Dis, vol.103, pp.1391-1396, 2019.

C. Carpino, L. Elvira-gonzález, L. Rubio, E. Peri, S. Davino et al., A comparative study of viral infectivity, accumulation and symptoms induced by broad bean wilt virus 1 isolates, J. Plant Pathol, vol.101, pp.275-281, 2019.

S. F. Elena and A. Fraile, García-Arenal, F. Evolution and emergence of plant viruses, Adv. Virus Res, vol.88, pp.161-191, 2014.

H. A. Hobbs, D. M. Eastburn, C. J. .;-d'arcy, J. D. Kindhart, J. B. Masiunas et al., Solanaceous weeds as possible sources of Cucumber mosaic virus in southern Illinois for aphid transmission to pepper, Plant Dis, vol.84, pp.1221-1224, 2000.

S. S. Barreto, M. Hallwass, O. M. Aquino, and A. K. Inoue-nagata, A study of weeds as potential inoculum sources for a tomato-infecting begomovirus in Central Brazil, Phytopathology, vol.103, pp.436-444, 2013.

S. Roselló, M. J. Díez, and F. Nuez, Viral diseases causing the greatest economic losses to the tomato crop. I. The Tomato spotted wilt virus-A review, Sci. Hortic, vol.67, pp.117-150, 1996.

H. Gong, C. Igiraneza, and L. Dusengemungu, Major In vitro techniques for potato virus elimination and post eradication detection methods. A review, Am. J. Potato Res, vol.96, pp.379-389, 2019.

S. García-andrés, D. M. Tomás, J. Navas-castillo, and E. Moriones, Resistant-driven selection of begomoviruses associated with the tomato yellow leaf curl disease, Virus Res, vol.146, pp.66-72, 2009.

C. López, J. Aramburu, L. Galipienso, S. Soler, F. Nuez et al., Evolutionary analysis of tomato Sw-5 resistance-breaking isolates of Tomato spotted wilt virus, J. Gen. Virol, vol.92, pp.210-215, 2011.

M. M. Goodin, D. Zaitlin, R. A. Naidu, and S. A. Lommel, Nicotiana benthamiana: Its history and future as a model for plant-pathogen interactions, © 2020 by the authors. Licensee MDPI, vol.21, pp.1015-1026, 2008.