, Management Authority to operate in Gonarezhou National Park and in Hwange National Park under permits

, )(c)(ii): 31, Gen, issue.1, p.23, 2007.

, 33/2011) and (ii) the South African National Parks to conduct our research in Kruger National Park. The project was conducted within the framework of the Research Platform "Production and Conservation in Partnership" (RP-PCP) and the Zone Atelier (LTSER) Hwange of the CNRS

, South African National Parks and Veterinary, 2009.

. Auth-o-r-co-ntr-i-b-uti-o-n,

, Elodie Wielgus: Conceptualization (equal); Data curation (equal)

, Formal analysis (lead)

, Writing-original draft (lead), Methodology (equal)

, Daniel Cornélis: Conceptualization (equal); Data curation (supporting)

, Methodology (supporting

, Resources (equal)

, Michel de Garine-Wichatitsky: Data curation (supporting)

, Writing-review & editing (supporting). Bradley Cain: Formal analysis (supporting)

, Funding acquisition (lead)

, Writing-review & editing (equal)

, Writing-review & editing (supporting)

, Eve Miguel: Data curation (supporting); Writing-review & editing (supporting). Hugo Valls-Fox: Data curation (supporting)

, Alexandre Caron: Conceptualization (equal); Data curation (equal)

, Methodology (equal

, Resources (equal)

, Writing-review & editing (equal), Simon Chamaillé-Jammes

, Conceptualization (equal)

, Data curation (equal); Formal analysis (lead)

, Methodology (equal

, Resources (equal); Writing-original draft (lead)

, Writing-review & editing (lead)

. Data-ava-i-l-a-b-i-l-i-t-y-s-tat-e-m-e-n-t-data,

O. Rci-d-elodie and . Wielgus,

E. A. Archie, C. J. Moss, and S. C. Alberts, The ties that bind: Genetic relatedness predicts the fission and fusion of social groups in wild African elephants, Proceedings of the Royal Society B: Biological Sciences, vol.273, pp.513-522, 2006.

T. W. Arnold, Uninformative parameters and model selection using Akaike's information criterion, The Journal of Wildlife Management, vol.74, pp.1175-1178, 2010.

F. Aureli, C. M. Schaffner, C. Boesch, S. K. Bearder, J. Call et al., Fission-fusion dynamics: New research frameworks, Current Anthropology, vol.49, pp.627-654, 2008.

A. L. Baden, T. H. Webster, and J. M. Kamilar, Resource seasonality and reproduction predict fission-fusion dynamics in black-and-white ruffed lemurs (Varecia variegata), American Journal of Primatology, vol.78, pp.256-279, 2016.

D. Bates, M. Mächler, B. Bolker, and S. Walker, Fitting linear mixed-effects models using lme4, Journal of Statistical Software, vol.67, pp.1-48, 2015.

H. Bauer and S. Van-der-merwe, Inventory of free-ranging lions Panthera leo in Africa, Oryx, vol.38, pp.26-31, 2004.

B. R. Beechler, A. E. Jolles, and V. O. Ezenwa, Evaluation of hematologic values in free-ranging African buffalo (Syncerus caffer), Journal of Wildlife Diseases, vol.45, pp.57-66, 2009.

S. Benhamou and D. Cornélis, Incorporating movement behavior and barriers to improve kernel home range space use estimates, Journal of Wildlife Management, vol.74, pp.1353-1360, 2010.

S. Benhamou, M. Valeix, S. Chamaillé-jammes, D. W. Macdonald, and A. J. Loveridge, Movement-based analysis of interactions in African lions, Animal Behavior, vol.90, pp.171-180, 2014.
URL : https://hal.archives-ouvertes.fr/hal-02107467

E. Bennitt, M. C. Bonyongo, and S. Harris, Habitat selection by African buffalo (Syncerus caffer) in response to landscape-level fluctuations in water availability on two temporal scales, PLoS One, vol.9, pp.1-14, 2014.

E. Bennitt, M. C. Bonyongo, and S. Harris, Cape buffalo (Syncerus caffer caffer) social dynamics in a flood-pulsed environment, Behavioral Ecology, vol.29, pp.93-105, 2018.

F. B. Bercovitch, P. S. Berry, R. G. Dwyer, J. M. Seddon, and A. W. Goldizen, Associations are more strongly correlated with space use than kinship in female eastern grey kangaroos, African Journal of Ecology, vol.48, pp.1-10, 2010.

E. C. Best, J. M. Seddon, R. G. Dwyer, and A. W. Goldizen, Social preference influences female community structure in a population of wild eastern grey kangaroos, Animal Behavior, vol.86, pp.1031-1040, 2013.

M. E. Brooks, K. Kristensen, K. J. Van-benthem, A. Magnusson, C. W. Berg et al., glmmTMB balances speed and flexibility among packages for zero-inflated generalized linear mixed modeling, The R Journal, vol.9, pp.378-400, 2017.

K. P. Burnham and D. R. Anderson, Model selection and multimodel inference: A practical information-theoretic approach, 2002.

C. Calenge, Exploring habitat selection by wildlife with adehabitat, Journal of Statistical Software, vol.22, pp.1-19, 2007.

A. Caron, P. C. Cross, and J. T. Du-toit, Ecological implications of bovine tuberculosis in African buffalo herds, Ecological Applications, vol.13, pp.1338-1345, 2003.

S. Chamaillé-jammes, H. Fritz, and F. Murindagomo, Spatial patterns of the NDVI-rainfall relationship at the seasonal and interannual time scales in an African savanna, International Journal of Remote Sensing, vol.27, pp.5185-5200, 2006.

S. Chamaillé-jammes, H. Fritz, and F. Murindagomo, Climatedriven fluctuations in surface-water availability and the buffering role of artificial pumping in an African savanna: Potential implication for herbivore dynamics, Austral Ecology, vol.32, pp.740-748, 2007.

C. A. Chapman, Ecological constraints on group size in three species of Neotropical primates, Folia Primatologica, vol.55, pp.1-9, 1990.

T. Clutton-brock, Mammal Societies, 2016.

R. C. Coles, P. C. Lee, and M. Talebi, Fission-fusion dynamics in Southern Muriquis (Brachyteles arachnoides) in continuous Brazilian Atlantic forest, International Journal of Primatology, vol.33, pp.93-114, 2012.

R. C. Connor, R. S. Wells, J. Mann, and A. J. Read, The bottlenose dolphin, Tursiops spp.: Social relationships in a fission-fusion society, Cetacean societies: Field studies of dolphins and whales, pp.91-126, 2000.

L. Conradt and T. J. Roper, Activity synchrony and social cohesion: A fission-fusion model, Proceedings of the Royal Society B-Biological Sciences, vol.267, pp.2213-2218, 2000.

D. Cornélis, S. Benhamou, G. Janeau, N. Morellet, M. Ouedraogo et al., Spatiotemporal dynamics of forage and water resources shape space use of West African savanna buffaloes, Journal of Mammalogy, vol.92, pp.1287-1297, 2011.

P. C. Cross, J. O. Lloyd-smith, and W. M. Getz, Disentangling association patterns in fission-fusion societies using African buffalo as an example, Animal Behavior, vol.69, pp.499-506, 2005.

M. De-garine-wichatitsky, A. Caron, C. Gomo, C. Foggin, K. Dutlow et al., Bovine tuberculosis in Buffaloes, Southern Africa. Emerging Infectious Diseases, vol.16, pp.884-885, 2010.

V. O. Ezenwa and A. E. Jolles, From host immunity to pathogen invasion: The effects of helminth coinfection on the dynamics of microparasites, Integrative and Comparative Biology, vol.51, pp.540-551, 2011.

S. M. Ferreira and P. J. Funston, Estimating lion population variables: Prey and disease effects in Kruger National Park, South Africa, Wildlife Research, vol.37, pp.194-206, 2010.

D. Fortin and M. E. Fortin, Group-size-dependent association between food profitability, predation risk and distribution of free-ranging bison, Animal Behavior, vol.78, pp.887-892, 2009.

D. Fortin, M. Fortin, H. L. Beyer, T. Duchesne, S. Courant et al., Group-size-mediated habitat selection and group fusion-fission dynamics of bison under predation risk, Ecology, vol.90, pp.2480-2490, 2009.

C. H. Frère, M. Krützen, J. Mann, J. J. Watson-capps, Y. J. Tsai et al., Home range overlap, matrilineal and biparental kinship drive female associations in bottlenose dolphins, Animal Behavior, vol.80, pp.481-486, 2010.

T. Furuichi, Factors underlying party size differences between chimpanzees and bonobos: A review and hypotheses for future study, Primates, vol.50, pp.197-209, 2009.

E. Gandiwa, I. M. Heitkönig, P. H. Eilers, and H. H. Prins, Rainfall variability and its impact on large mammal populations in a complex of semi-arid African savanna protected areas, Tropical Ecology, vol.57, pp.163-180, 2016.

E. Gandiwa and P. Zisadza, Wildlife management in Gonarezhou National Park, Southeast Zimbabwe: Climate change and implications for management, Nature and Faune, vol.25, pp.96-105, 2010.

W. P. Gertenbach, Rainfall patterns in the Kruger National Park, Koedoe, vol.23, pp.35-43, 1980.

W. P. Gertenbach, Landscapes of the Kruger National Park, Koedoe, vol.26, pp.9-121, 1983.

R. Groom, Carnivore Densities in Gonarezhou National Park Results of the May, Spoor Survey. African Wildlife Conservation Fund, 2009.

D. J. Halley, M. E. Vandewalle, M. Mari, and C. Taolo, Herdswitching and long-distance dispersal in female African buffalo Syncerus caffer, African Journal of Ecology, vol.40, pp.97-99, 2002.

R. A. Hill and P. C. Lee, Predation risk as an influence on group size in cercopithecoid primates: Implications for social structure, Journal of Zoology London, vol.245, pp.447-456, 1998.

K. Isvaran, Intraspecific variation in group size in the blackbuck antelope: The roles of habitat structure and forage at different spatial scales, Oecologia, vol.154, pp.435-444, 2007.

P. J. Jarman, The social organisation of Antelope in relation to their ecology, Behaviour, vol.48, pp.215-267, 1974.

K. Kashima, H. Ohtsuki, and A. Satake, Fission-fusion bat behavior as a strategy for balancing the conflicting needs of maximizing information accuracy and minimizing infection risk, Journal of Theoretical Biology, vol.318, pp.101-109, 2013.

J. L. Kelley, L. J. Morrell, C. Inskip, J. Krause, and D. P. Croft, Predation risk shapes social networks in fission-fusion populations, PLoS One, vol.6, 2011.

G. Kerth and B. König, Fission, fusion and nonrandom associations in female Bechstein'S Bats (Myotis Bechsteinii), Behaviour, vol.136, p.1711, 1999.

M. La-grange, Chemical and Physical Restraint of Wild Animals-A Training Field Manual for African Species, Zimbabwe Veterinary Association Wildlife Group and International Wildlife Veterinary Services, pp.250-259, 2006.

J. Lehmann and C. Boesch, To fission or to fusion: Effects of community size on wild chimpanzee (Pan troglodytes verus) social organisation, Behavioral Ecology and Sociobiology, vol.56, pp.207-216, 2004.

F. Lesmerises, C. J. Johnson, and M. St-laurent, Landscape knowledge is an important driver of the fission dynamics of an alpine ungulate, Animal Behavior, vol.140, pp.39-47, 2018.

M. Loretto, R. Schuster, C. Itty, P. Marchand, F. Genero et al., Fission-fusion dynamics over large distances in raven non-breeders, Scientific Reports, vol.7, 2017.

A. J. Loveridge, A. W. Searle, F. Murindagomo, and D. W. Macdonald, The impact of sport-hunting on the population dynamics of an African lion population in a protected area, Biological Conservation, vol.134, pp.548-558, 2007.

D. Lüdecke, D. Makowski, and P. Waggoner, Performance: Assessment of Regression Models Performance, 2019.

A. Megaze, M. Balakrishnan, and G. Belay, Diurnal activity budget of African buffalo (Syncerus caffer Sparrman, 1779) in Chebera Churchura National Park, Ethiopia. African Journal of Ecology, vol.56, pp.436-444, 2016.

S. Nakagawa, P. C. Johnson, and H. Schielzeth, The coefficient of determination R 2 and intra-class correlation coefficient from generalized linear mixed-effects models revisited and expanded, Journal of the Royal Society, Interface, vol.14, 2017.

G. J. Parra, P. J. Corkeron, and P. Arnold, Grouping and fission-fusion dynamics in Australian snubfin and Indo-Pacific humpback dolphins, Animal Behavior, vol.82, pp.1423-1433, 2011.

O. Pays, S. Benhamou, R. Helder, and J. Gerard, The dynamics of group formation in large mammalian herbivores: An analysis in the European roe deer, Animal Behavior, vol.74, pp.1429-1441, 2007.
URL : https://hal.archives-ouvertes.fr/hal-01503332

D. Pépin and J. Gerard, Group dynamics and local population density dependence of group size in the Pyrenean chamois, Rupicapra pyrenaica, Animal Behavior, vol.75, pp.361-369, 2008.

B. Pinacho-guendulain and G. Ramos-fernández, Influence of fruit availability on the fission-fusion dynamics of spider monkeys (Ateles geoffroyi), International Journal of Primatology, vol.38, pp.1-19, 2017.

L. Polansky, G. Wittemyer, P. C. Cross, C. J. Tambling, and W. M. Getz, From moonlight to movement and synchronized randomness: Fourier and wavelet analyses of animal location time series data, Ecology, vol.91, pp.1506-1518, 2010.

H. H. Prins, Buffalo herd structure and its repercussions for condition of individual African buffalo cows, Ethology, vol.81, pp.47-71, 1989.

H. H. Prins, Ecology and behaviour of the African buffalo: Social inequality and decision making, 1996.

A. Purdon and R. J. Van-aarde, Water provisioning in Kruger National Park alters elephant spatial utilisation patterns, Journal of Arid Environments, vol.141, pp.45-51, 2017.

. R-core-team, R: A language and environment for statistical computing, 2016.

S. J. Ryan, C. U. Knechtel, and W. M. Getz, Range and habitat selection of African buffalo in South Africa, Journal of Wildlife Management, vol.70, pp.764-776, 2006.

Y. Shimooka, Seasonal variation in association patterns of wild spider monkeys (Ateles belzebuth belzebuth) at La Macarena, Colombia, Primates, vol.44, pp.83-90, 2003.

A. R. Sinclair, The African buffalo: A study of resource limitation of populations, 1977.

C. Sueur, A. J. King, L. Conradt, G. Kerth, D. Lusseau et al., Collective decision-making and fission-fusion dynamics: A conceptual framework, Oikos, vol.120, issue.11, pp.1608-1617, 2011.

C. J. Tambling, D. J. Druce, M. W. Hayward, J. G. Castley, J. Adendorff et al., Spatial and temporal changes in group dynamics and range use enable anti-predator responses in African buffalo, Ecology, vol.93, pp.1297-1304, 2012.

H. Valls-fox, S. Chamaillé-jammes, M. De-garine-wichatitsky, A. Perrotton, N. Courbin et al., Water and cattle shape habitat selection by wild herbivores at the edge of a protected area, Animal Conservation, vol.21, pp.365-375, 2018.
URL : https://hal.archives-ouvertes.fr/hal-01960150

S. N. Wood, Fast stable restricted maximum likelihood and marginal likelihood estimation of semiparametric generalized linear models, Journal of the Royal Statistical Society: Series B (Statistical Methodology), vol.73, pp.3-36, 2011.

S. N. Wood, Generalized Additive Models: An Introduction with R, 2017.

M. Zvidzai, A. Murwira, A. Caron, and M. De-garine-wichatitsky, Waterhole use patterns at the wildlife/livestock interface in a semiarid savanna of Southern Africa, International Journal of Development and Sustainability, vol.2, pp.455-471, 2013.