T. G. Gilmanov, J. F. Soussana, L. Aires, V. Allard, C. Ammann et al., Partitioning European grassland net ecosystem CO 2 exchange into gross primary productivity and ecosystem respiration using light response function analysis, Agric. Ecosyst. Environ, vol.121, pp.93-120, 2007.
URL : https://hal.archives-ouvertes.fr/hal-02664219

V. O. Snow, C. A. Rotz, A. D. Moore, R. Martin-clouaire, I. R. Johnson et al., The challenges and some solutions to process-based modelling of grazed agricultural systems, Environ. Model. Softw, vol.62, pp.420-436, 2014.
URL : https://hal.archives-ouvertes.fr/hal-02635654

M. Fischer, O. Bossdorf, S. Gockel, F. Hänsel, A. Hemp et al., Implementing large-scale and long-term functional biodiversity research: The Biodiversity Exploratories, Basic Appl. Ecol, vol.11, pp.473-485, 2010.

A. Tribot, J. Deter, and N. Mouquet, Integrating the aesthetic value of landscapes and biological diversity, Proc. Biol. Sci, vol.285, 2018.
URL : https://hal.archives-ouvertes.fr/hal-02002248

, Agronomy 2020, vol.10, pp.259-278

D. Tilman, J. Hill, and C. Lehman, Carbon-negative biofuels from low-input high-diversity grassland biomass, Science, vol.314, pp.1598-1600, 2006.

P. Dass, B. Z. Houlton, Y. Wang, and D. Warlind, Grasslands may be more reliable carbon sinks than forests in California, Environ. Res. Lett, vol.13, p.74027, 2018.

C. Carvalho-santos, A. T. Monteiro, S. Arenas-castro, F. Greifeneder, B. Marcos et al., Ecosystem services in a protected mountain range of Portugal: Satellite-based products for state and trend analysis, Remote Sens, vol.10, 1573.

P. E. Levy, D. C. Mobbs, S. K. Jones, R. Milne, C. Campbell et al., Simulation of fluxes of greenhouse gases from European grasslands using the DNDC model, Agric. Ecosyst. Environ, vol.121, pp.186-192, 2007.

J. Chang, P. Ciais, N. Viovy, N. Vuichard, B. Sultan et al., The greenhouse gas balance of European grasslands, Glob. Chang. Biol, vol.21, pp.3748-3761, 2015.
URL : https://hal.archives-ouvertes.fr/hal-01196330

C. Oertel, J. Matschullat, K. Zurba, F. Zimmermann, and S. Erasmi, Greenhouse gas emissions from soils-A review, Geochemistry, vol.76, pp.327-352, 2016.

O. Bastian, The role of biodiversity in supporting ecosystem services in Natura 2000 sites, Ecol. Indic, vol.24, pp.12-22, 2013.

L. A. Turnbull, F. Isbell, D. W. Purves, M. Loreau, and A. Hector, Understanding the value of plant diversity for ecosystem functioning through niche theory, Proc. R. Soc. B Biol. Sci, vol.283, 2016.
URL : https://hal.archives-ouvertes.fr/hal-02962585

S. Plantureux, A. Peeters, and D. Mccracken, Biodiversity in intensive grasslands: Effect of management, improvement and challenges, Agron. Res, vol.3, pp.153-164, 2005.
URL : https://hal.archives-ouvertes.fr/hal-02677402

R. Van-klink, F. Van-der-plas, C. G. Van-noordwijk, M. F. Wallisdevries, and H. Olff, Effects of large herbivores on grassland arthropod diversity, Biol. Rev. Camb. Philos. Soc, vol.90, pp.347-366, 2015.

M. Lange, N. Eisenhauer, C. A. Sierra, H. Bessler, C. Engels et al., Plant diversity increases soil microbial activity and soil carbon storage, Nat. Commun, vol.6, pp.1-8, 2015.
URL : https://hal.archives-ouvertes.fr/hal-02475329

S. Yachi and M. Loreau, Biodiversity and ecosystem productivity in a fluctuating environment: The insurance hypothesis, Proc. Natl. Acad. Sci, vol.96, pp.1463-1468, 1999.

J. Norberg, D. P. Swaney, J. Dushoff, J. Lin, R. Casagrandi et al., Phenotypic diversity and ecosystem functioning in changing environments: A theoretical framework, Proc. Natl. Acad. Sci, vol.98, pp.11376-11381, 2001.

A. Thébault, P. Mariotte, C. J. Lortie, and A. S. Macdougall, Land management trumps the effects of climate change and elevated CO2 on grassland functioning, J. Ecol, vol.102, pp.896-904, 2014.

A. Vogel, M. Scherer-lorenzen, and A. Weigelt, Grassland resistance and resilience after drought depends on management intensity and species richness, PLoS ONE, vol.7, 2012.

D. Craven, F. Isbell, P. Manning, J. Connolly, H. Bruelheide et al., Plant diversity effects on grassland productivity are robust to both nutrient enrichment and drought, Philos. Trans. R. Soc. B Biol. Sci, vol.371, 2016.
URL : https://hal.archives-ouvertes.fr/hal-02962385

E. E. Cleland, Biodiversity and ecosystem stability, Nat. Educ. Knowl, 2011.

J. M. Craine, T. W. Ocheltree, J. B. Nippert, E. G. Towne, A. M. Skibbe et al., Global diversity of drought tolerance and grassland climate-change resilience, Nat. Clim. Chang, vol.3, pp.63-67, 2013.

F. Volaire, K. Barkaoui, and M. Norton, Designing resilient and sustainable grasslands for a drier future: Adaptive strategies, functional traits and biotic interactions, Eur. J. Agron, vol.52, pp.81-89, 2014.
URL : https://hal.archives-ouvertes.fr/hal-02804811

J. M. Cowles, P. D. Wragg, A. J. Wright, J. S. Powers, and D. Tilman, Shifting grassland plant community structure drives positive interactive effects of warming and diversity on aboveground net primary productivity, Glob. Chang. Biol, vol.22, pp.741-749, 2016.

A. J. Wright, H. De-kroon, E. J. Visser, T. Buchmann, A. Ebeling et al., Plants are less negatively affected by flooding when growing in species-rich plant communities, New Phytol, vol.213, pp.645-656, 2017.

U. Zeller, N. Starik, and T. Göttert, Biodiversity, land use and ecosystem services: An organismic and comparative approach to different geographical regions, Glob. Ecol. Conserv, vol.10, p.25, 2017.

M. Loreau, C. Mazancourt, and . De, Biodiversity and ecosystem stability: A synthesis of underlying mechanisms, Ecol. Lett, vol.16, pp.106-115, 2013.

J. H. Spangenberg, C. Görg, D. T. Truong, V. Tekken, J. V. Bustamante et al., Provision of ecosystem services is determined by human agency, not ecosystem functions. Four case studies, Int. J. Biodivers. Sci. Ecosyst. Serv. Manag, vol.10, pp.40-53, 2014.

L. Da-silveira-pontes, V. Maire, J. Schellberg, and F. Louault, Grass strategies and grassland community responses to environmental drivers: A review, Agron. Sustain. Dev, vol.35, pp.1297-1318, 2015.
URL : https://hal.archives-ouvertes.fr/hal-01325121

E. Gaujour, B. Amiaud, C. Mignolet, and S. Plantureux, Factors and processes affecting plant biodiversity in permanent grasslands. A review, Agron. Sustain. Dev, vol.32, pp.133-160, 2012.
URL : https://hal.archives-ouvertes.fr/hal-00930482

W. Thuiller, C. Albert, M. B. Araújo, P. M. Berry, M. Cabeza et al., Predicting global change impacts on plant species' distributions: Future challenges, Perspect. Plant Ecol. Evol. Syst, vol.9, pp.137-152, 2008.
URL : https://hal.archives-ouvertes.fr/halsde-00281677

G. Balent, D. Alard, V. Blanfort, and I. Poudevigne, Pratiques de gestion, biodiversité floristique et durabilité des prairies, vol.160, pp.385-402, 1999.

M. Van-oijen, G. Bellocchi, and M. Höglind, Effects of climate change on grassland biodiversity and productivity: The need for a diversity of models, vol.8, p.14, 2018.
URL : https://hal.archives-ouvertes.fr/hal-01717916

C. Porqueddu, S. Ates, M. Louhaichi, A. P. Kyriazopoulos, G. Moreno et al., Mediterranean-climate zones: Past trends, current status and future research priorities, Grass Forage Sci, vol.71, pp.1-35, 2016.

M. Rapacz, Å. Ergon, M. Höglind, M. Jørgensen, B. Jurczyk et al., Overwintering of herbaceous plants in a changing climate. Still more questions than answers, Plant Sci, vol.225, pp.34-44, 2014.

L. Rhazi, P. Grillas, E. Saber, M. Rhazi, L. Brendonck et al., Vegetation of Mediterranean temporary pools: A fading jewel? Hydrobiologia, vol.689, pp.23-36, 2012.

L. Ma, J. D. Derner, R. D. Harmel, J. Tatarko, A. D. Moore et al., Application of grazing land models in ecosystem management: Current status and next frontiers, Advances in Agronomy, p.0065211319300719, 2019.
URL : https://hal.archives-ouvertes.fr/hal-02509527

S. Burri, P. A. Niklaus, K. Grassow, N. Buchmann, and A. Kahmen, Effects of plant productivity and species richness on the drought response of soil respiration in temperate grasslands, PLoS ONE, vol.13, 2018.

F. Isbell, D. Craven, J. Connolly, M. Loreau, B. Schmid et al., Biodiversity increases the resistance of ecosystem productivity to climate extremes, Nature, vol.526, pp.574-577, 2015.
URL : https://hal.archives-ouvertes.fr/hal-02959854

D. Boeck, H. J. Bloor, J. M. Kreyling, J. Ransijn, J. C. Nijs et al., Patterns and drivers of biodiversity-stability relationships under climate extremes, J. Ecol, vol.106, pp.890-902, 2018.
URL : https://hal.archives-ouvertes.fr/hal-02625633

A. Hector, B. Schmid, C. Beierkuhnlein, M. C. Caldeira, M. Diemer et al., Plant diversity and productivity experiments in European grasslands, Science, vol.286, pp.1123-1127, 1999.

A. I. Roca-fernández, J. L. Peyraud, L. Delaby, and R. Delagarde, Pasture intake and milk production of dairy cows rotationally grazing on multi-species swards, Animal, vol.10, pp.1448-1456, 2016.

J. Connolly, M. Sebastià, L. Kirwan, J. A. Finn, R. Llurba et al., Weed suppression greatly increased by plant diversity in intensively managed grasslands: A continental-scale experiment, J. Appl. Ecol, vol.55, pp.852-862, 2018.
URL : https://hal.archives-ouvertes.fr/hal-02623722

K. A. Orford, P. J. Murray, I. P. Vaughan, and J. Memmott, Modest enhancements to conventional grassland diversity improve the provision of pollination services, J. Appl. Ecol, vol.53, pp.906-915, 2016.

A. Lüscher, I. Mueller-harvey, J. F. Soussana, R. M. Rees, and J. L. Peyraud, Potential of legume-based grasslandLivestock systems in Europe: A review. Grass Forage Sci, vol.69, pp.206-228, 2014.

T. E. Crews and B. E. Rumsey, What agriculture can learn from native ecosystems in building soil organic matter: A review, vol.9, 2017.

H. Chen, L. Mommer, J. Van-ruijven, H. De-kroon, C. Fischer et al., Plant species richness negatively affects root decomposition in grasslands, J. Ecol, vol.105, pp.209-218, 2017.

, Agronomy 2020, vol.10, pp.259-280

N. Kagiya, T. Reinsch, F. Taube, J. Salminen, C. Kluß et al., Turnover rates of roots vary considerably across temperate forage species, Soil Biol. Biochem, vol.139, 2019.

C. Chen, H. Y. Chen, X. Chen, and Z. Huang, Meta-analysis shows positive effects of plant diversity on microbial biomass and respiration, Nat. Commun, vol.10, pp.1-10, 2019.

W. W. Weisser, C. Roscher, S. T. Meyer, A. Ebeling, G. Luo et al., Biodiversity effects on ecosystem functioning in a 15-year grassland experiment: Patterns, mechanisms, and open questions, Basic Appl. Ecol, vol.23, pp.1-73, 2017.
URL : https://hal.archives-ouvertes.fr/hal-02480455

Y. Yang, D. Tilman, G. Furey, and C. Lehman, Soil carbon sequestration accelerated by restoration of grassland biodiversity, Nat. Commun, vol.10, pp.1-7, 2019.

X. Wei, P. B. Reich, and S. E. Hobbie, Legumes regulate grassland soil N cycling and its response to variation in species diversity and N supply but not CO2, Glob. Chang. Biol, vol.25, pp.2396-2409, 2019.

E. M. Elst, H. J. De-boeck, L. Vanmaele, M. Verlinden, and P. Dhliwayo, Nijs, I. Impact of climate extremes modulated by species characteristics and richness, Perspect. Plant Ecol. Evol. Syst, vol.24, pp.80-92, 2017.

D. S. Gruner, M. E. Bracken, S. A. Berger, B. K. Eriksson, L. Gamfeldt et al., Effects of experimental warming on biodiversity depend on ecosystem type and local species composition, vol.126, pp.8-17, 2016.

S. P. Harrison, E. S. Gornish, and S. Copeland, Climate-driven diversity loss in a grassland community, Proc. Natl. Acad. Sci, vol.112, pp.8672-8677, 2015.

J. P. Grime, J. D. Fridley, A. P. Askew, K. Thompson, J. G. Hodgson et al., Long-term resistance to simulated climate change in an infertile grassland, Proc. Natl. Acad. Sci, vol.105, pp.10028-10032, 2008.

S. Bagella, L. Salis, G. M. Marrosu, I. Rossetti, S. Fanni et al., Effects of long-term management practices on grassland plant assemblages in Mediterranean cork oak silvo-pastoral systems, Plant Ecol, vol.214, pp.621-631, 2013.

G. Seddaiu, S. Bagella, A. Pulina, C. Cappai, L. Salis et al., Mediterranean cork oak wooded grasslands: Synergies and trade-offs between plant diversity, pasture production and soil carbon, Agroforest. Syst, vol.92, pp.893-908, 2018.

N. Demalach, E. Zaady, and R. Kadmon, Contrasting effects of water and nutrient additions on grassland communities: A global meta-analysis, Glob. Ecol. Biogeogr, vol.26, pp.983-992, 2017.

J. Humbert, J. M. Dwyer, A. Andrey, and R. Arlettaz, Impacts of nitrogen addition on plant biodiversity in mountain grasslands depend on dose, application duration and climate: A systematic review, Glob. Chang. Biol, vol.22, pp.110-120, 2016.

M. B. Soons, M. M. Hefting, E. Dorland, L. P. Lamers, C. Versteeg et al., Nitrogen effects on plant species richness in herbaceous communities are more widespread and stronger than those of phosphorus, Biol. Conserv, vol.212, pp.390-397, 2017.

M. Tälle, B. Deák, P. Poschlod, O. Valkó, L. Westerberg et al., Grazing vs. Mowing: A meta-analysis of biodiversity benefits for grassland management, Agric. Ecosyst. Environ, vol.222, pp.200-212, 2016.

L. Johansen, S. Taugourdeau, K. A. Hovstad, and S. Wehn, Ceased grazing management changes the ecosystem services of semi-natural grasslands, Ecosyst. People, vol.15, pp.192-203, 2019.

P. Török, K. Penksza, E. Tóth, A. Kelemen, J. Sonkoly et al., Vegetation type and grazing intensity jointly shape grazing effects on grassland biodiversity, Ecol. Evol, vol.8, pp.10326-10335, 2018.

D. Tilman, P. B. Reich, and F. Isbell, Biodiversity impacts ecosystem productivity as much as resources, disturbance, or herbivory, Proc. Natl. Acad. Sci, vol.109, pp.10394-10397, 2012.

Z. Y. Yuan, F. Jiao, Y. H. Li, and R. L. Kallenbach, Anthropogenic disturbances are key to maintaining the biodiversity of grasslands, Sci. Rep, vol.6, pp.1-8, 2016.

S. A. Grant, L. Torvell, E. M. Sim, J. L. Small, and R. H. Armstrong, Controlled grazing studies on Nardus grassland: Effects of between-tussock sward height and species of grazer on Nardus utilization and floristic composition in two fields in Scotland, J. Appl. Ecol, vol.33, pp.1053-1064, 1996.

T. Lunnan, J. Todnem, and M. Jørgensen, Botanical composition of grassland for silage in mountain districts of Norway, Proceedings of the 27th General Meeting of the European Grassland Federation, pp.274-276, 2018.

M. Loreau and A. Hector, Partitioning selection and complementarity in biodiversity experiments, Nature, vol.412, pp.72-76, 2001.

B. J. Cardinale, K. L. Matulich, D. U. Hooper, J. E. Byrnes, E. Duffy et al., The functional role of producer diversity in ecosystems, Am. J. Bot, vol.98, pp.572-592, 2011.

S. Da, L. Pontes, V. Maire, F. Louault, J. Soussana et al., Impacts of species interactions on grass community productivity under contrasting management regimes, Oecologia, vol.168, pp.761-771, 2012.
URL : https://hal.archives-ouvertes.fr/hal-02651104

J. Ingrisch, S. Karlowsky, A. Anadon-rosell, R. Hasibeder, A. König et al., Land use alters the drought responses of productivity and CO 2 fluxes in mountain grassland, Ecosystems, vol.21, pp.689-703, 2018.

M. C. Rillig, M. Ryo, A. Lehmann, C. A. Aguilar-trigueros, S. Buchert et al., The role of multiple global change factors in driving soil functions and microbial biodiversity, Science, vol.366, pp.886-890, 2019.

J. Liu, X. Li, Q. Ma, X. Zhang, Y. Chen et al., Nitrogen addition reduced ecosystem stability regardless of its impacts on plant diversity, J. Ecol, vol.107, pp.2427-2435, 2019.

M. L. Rosenzweig, Species Diversity in Space and Time, 1995.

J. B. Grace, T. Michael-anderson, M. D. Smith, E. Seabloom, S. J. Andelman et al., Does species diversity limit productivity in natural grassland communities?, Ecol. Lett, vol.10, pp.680-689, 2007.

G. F. Veen, W. H. Van-der-putten, and T. M. Bezemer, Biodiversity-ecosystem functioning relationships in a long-term non-weeded field experiment, Ecology, vol.99, pp.1836-1846, 2018.

P. Kardol, N. Fanin, and D. A. Wardle, Long-term effects of species loss on community properties across contrasting ecosystems, Nature, vol.557, pp.710-713, 2018.
URL : https://hal.archives-ouvertes.fr/hal-02626980

L. Brilli, L. Bechini, M. Bindi, M. Carozzi, D. Cavalli et al., Review and analysis of strengths and weaknesses of agro-ecosystem models for simulating C and N fluxes, Sci. Total Environ, vol.598, pp.445-470, 2017.
URL : https://hal.archives-ouvertes.fr/hal-01537755

P. Korhonen, T. Palosuo, T. Persson, M. Höglind, G. Jégo et al., Modelling grass yields in northern climates a comparison of three growth models for timothy, Field Crops Res, vol.224, pp.37-47, 2018.

T. Persson, M. Höglind, M. Van-oijen, P. Korhonen, T. Palosuo et al., Simulation of timothy nutritive value: A comparison of three process-based models, Field Crops Res, vol.231, pp.81-92, 2019.

L. Strullu, N. Beaudoin, P. Thiébeau, B. Julier, B. Mary et al., Simulation using the STICS model of C&N dynamics in alfalfa from sowing to crop destruction, Eur. J. Agron, vol.2020, p.125948
URL : https://hal.archives-ouvertes.fr/hal-02622466

P. Calanca, Modelling the impacts of seasonal drought on herbage growth under climate change, Adv. Anim. Biosci, vol.7, pp.231-232, 2016.

M. Jouven, P. Carrère, and R. Baumont, Model predicting dynamics of biomass, structure and digestibility of herbage in managed permanent pastures. 1. Model description. Grass Forage Sci, vol.61, pp.112-124, 2006.
URL : https://hal.archives-ouvertes.fr/hal-02653991

N. Fitton, M. Bindi, L. Brilli, R. Cichota, C. Dibari et al., Modelling biological N fixation and grass-legume dynamics with process-based biogeochemical models of varying complexity, Eur. J. Agron, vol.106, pp.58-66, 2019.
URL : https://hal.archives-ouvertes.fr/hal-02166488

F. Gillet, Modelling vegetation dynamics in heterogeneous pasture-woodland landscapes, Ecol. Model, vol.217, pp.1-18, 2008.
URL : https://hal.archives-ouvertes.fr/hal-00357747

D. Tilman, Community invasibility, recruitment limitation, and grassland biodiversity, Ecology, vol.78, pp.81-92, 1997.

D. Tilman, Resources: A graphical-mechanistic approach to competition and predation, Am. Nat, vol.116, pp.362-393, 1980.

G. Hardin, The competitive exclusion principle, Science, vol.131, pp.1292-1297, 1960.

T. Moulin, A. Perasso, and F. Gillet, Modelling vegetation dynamics in managed grasslands: Responses to drivers depend on species richness, Ecol. Model, vol.374, pp.22-36, 2018.
URL : https://hal.archives-ouvertes.fr/hal-01779748

S. Schwinning and A. J. Parsons, Interactions between grasses and legumes: Understanding variability in species composition, Legumes in Sustainable Farming Systems, British Grassland Society Occasional Symposium, 1996.

D. Younie and . Ed, , pp.153-163, 1996.

M. Loreau, N. Mouquet, and A. Gonzalez, Biodiversity as spatial insurance in heterogeneous landscapes, Proc. Natl. Acad. Sci, vol.100, pp.12765-12770, 2003.

C. De-mazancourt, F. Isbell, A. Larocque, F. Berendse, E. De-luca et al., Predicting ecosystem stability from community composition and biodiversity, Ecol. Lett, vol.16, pp.617-625, 2013.
URL : https://hal.archives-ouvertes.fr/hal-02331250

S. Siehoff, G. Lennartz, I. C. Heilburg, M. Roß-nickoll, H. T. Ratte et al., Process-based modeling of grassland dynamics built on ecological indicator values for land use, Ecol. Model, vol.222, pp.3854-3868, 2011.

M. Westoby, B. Walker, and I. Noy-meir, Opportunistic management for rangelands not at equilibrium. J. Range Manag, vol.42, pp.266-274, 1989.

F. Taubert, K. Frank, and A. Huth, A review of grassland models in the biofuel context, Ecol. Model, vol.245, pp.84-93, 2012.

L. Weiss, H. Pfestorf, F. May, K. Körner, S. Boch et al., Grazing response patterns indicate isolation of semi-natural European grasslands, vol.123, pp.599-612, 2014.

F. May, V. Grimm, and F. Jeltsch, Reversed effects of grazing on plant diversity: The role of below-ground competition and size symmetry, Oikos, vol.118, pp.1830-1843, 2009.

D. G. Milchunas, O. E. Sala, and W. Lauenroth, A generalized model of the effects of grazing by large herbivores on grassland community structure, Am. Nat, vol.132, pp.87-106, 1988.

J. W. Fox, The intermediate disturbance hypothesis should be abandoned, Trends Ecol. Evol, vol.28, pp.86-92, 2013.

D. Lohmann, T. Guo, and B. Tietjen, Zooming in on coarse plant functional types: Simulated response of savanna vegetation composition in response to aridity and grazing, Theor. Ecol, vol.11, pp.161-173, 2018.

A. K. Bittebiere, C. Mony, B. Clément, and M. Garbey, Modeling competition between plants using an Individual Based Model: Methods and effects on the growth of two species with contrasted growth forms, Ecol. Model, vol.234, pp.38-50, 2012.
URL : https://hal.archives-ouvertes.fr/hal-00716031

P. M. Van-bodegom, J. C. Douma, and L. M. Verheijen, A fully traits-based approach to modeling global vegetation distribution, Proc. Natl. Acad. Sci, vol.111, pp.13733-13738, 2014.

D. C. Laughlin, C. Joshi, P. M. Van-bodegom, Z. A. Bastow, and P. Z. Fulé, A predictive model of community assembly that incorporates intraspecific trait variation, Ecol. Lett, vol.15, pp.1291-1299, 2012.

D. C. Laughlin, C. Joshi, S. J. Richardson, D. A. Peltzer, N. W. Mason et al., Quantifying multimodal trait distributions improves trait-based predictions of species abundances and functional diversity, J. Veg. Sci, vol.26, pp.46-57, 2015.

M. Van-oijen and M. Höglind, Toward a Bayesian procedure for using process-based models in plant breeding, with application to ideotype design, Euphytica, vol.207, pp.627-643, 2016.

J. Dole?al, V. Lanta, O. Mudrák, and J. Lep?, Seasonality promotes grassland diversity: Interactions with mowing, fertilization and removal of dominant species, J. Ecol, vol.107, pp.203-215, 2019.

G. Xenakis, D. Ray, and M. Mencuccini, Sensitivity and uncertainty analysis from a coupled 3-PG and soil organic matter decomposition model, Ecol. Model, vol.219, pp.1-16, 2008.

J. Thornley, Simulating grass-legume dynamics: A phenomenological submodel, Ann. Bot, vol.88, pp.905-913, 2001.

R. Confalonieri and . Cosmo, A simple approach for reproducing plant community dynamics using a single instance of generic crop simulators, Ecol. Model, vol.286, pp.1-10, 2014.

E. Movedi, G. Bellocchi, G. Argenti, L. Paleari, F. Vesely et al., Development of generic crop models for simulation of multi-species plant communities in mown grasslands, Ecol. Model, vol.401, pp.111-128, 2019.
URL : https://hal.archives-ouvertes.fr/hal-02154677

E. Malézieux, Y. Crozat, C. Dupraz, M. Laurans, D. Makowski et al., Valantin-Morison, M. Mixing plant species in cropping systems: Concepts, tools and models. A review, Agron. Sustain. Dev, vol.29, pp.43-62, 2009.

, Agronomy 2020, vol.10, pp.259-283

B. Sakschewski, W. Von-bloh, A. Boit, A. Rammig, J. Kattge et al., Leaf and stem economics spectra drive diversity of functional plant traits in a dynamic global vegetation model, Glob. Chang. Biol, vol.21, pp.2711-2725, 2015.

G. Louarn and L. Faverjon, A generic individual-based model to simulate morphogenesis, CN acquisition and population dynamics in contrasting forage legumes, Ann. Bot, vol.121, pp.875-896, 2018.
URL : https://hal.archives-ouvertes.fr/hal-02622343

N. Colbach, A. Gardarin, and D. Moreau, The response of weed and crop species to shading: Which parameters explain weed impacts on crop production? Field Crops Res, vol.238, pp.45-55, 2019.

, Modelling Crop-Weed Interactions

M. Kropff and . Van-laar, , 1993.

M. Van-noordwijk and B. Lusiana, WaNuLCAS, a model of water, nutrient and light capture in agroforestry systems, Agrofor. Syst, vol.43, pp.217-242, 1998.

M. Van-oijen, J. Dauzat, J. Harmand, G. Lawson, and P. Vaast, Coffee agroforestry systems in Central America: II. Development of a simple process-based model and preliminary results, Agrofor. Syst, vol.80, pp.361-378, 2010.

G. Corre-hellou, M. Faure, M. Launay, N. Brisson, and Y. Crozat, Adaptation of the STICS intercrop model to simulate crop growth and N accumulation in pea-barley intercrops, Field Crops Res, vol.113, pp.72-81, 2009.
URL : https://hal.archives-ouvertes.fr/hal-02665845

P. Lazzarotto, P. Calanca, and J. Fuhrer, Dynamics of grass clover mixtures: An analysis of the response to management with the PROductive GRASsland Simulator (PROGRASS), Ecol. Model, vol.220, pp.703-724, 2009.

J. M. Sharp, G. R. Edwards, and M. J. Jeger, A spatially explicit population model of the effect of spatial scale of heterogeneity in grass-clover grazing systems, J. Agric. Sci, vol.152, pp.394-407, 2014.

M. Biondini, A three-dimensional spatial model for plant competition in a heterogeneous soil environment, Ecol. Model, vol.142, pp.189-225, 2001.

J. Vos, J. B. Evers, G. H. Buck-sorlin, B. Andrieu, M. Chelle et al., Functional-structural plant modelling: A new versatile tool in crop science, J. Exp. Bot, vol.61, pp.2101-2115, 2010.
URL : https://hal.archives-ouvertes.fr/hal-01132296

L. Faverjon, A. Escobar-gutiérrez, I. Litrico, B. Julier, and G. Louarn, A generic individual-based model can predict yield, nitrogen content, and species abundance in experimental grassland communities, J. Exp. Bot, vol.70, pp.2491-2504, 2019.
URL : https://hal.archives-ouvertes.fr/hal-02622121

J. Soussana, V. Maire, N. Gross, B. Bachelet, L. Pagès et al., A grassland model simulating the role of plant traits for community dynamics and ecosystem functioning. Parameterization and evaluation, Ecol. Model, vol.231, pp.134-145, 2012.
URL : https://hal.archives-ouvertes.fr/hal-00707633

V. Maire, J. F. Soussana, N. Gross, B. Bachelet, L. Pagès et al., Plasticity of plant form and function sustains productivity and dominance along environment and competition gradients. A modeling experiment with Gemini, Ecol. Model, vol.254, pp.80-91, 2013.
URL : https://hal.archives-ouvertes.fr/hal-00706761

I. J. Wright, P. B. Reich, M. Westoby, D. D. Ackerly, Z. Baruch et al., The worldwide leaf economics spectrum, Nature, vol.428, pp.821-827, 2004.

V. Maire, N. Gross, D. Hill, R. Martin, C. Wirth et al., Disentangling coordination among functional traits using an individual-centred model: Impact on plant performance at intra-and inter-specific levels, PLoS ONE, vol.8, 2013.
URL : https://hal.archives-ouvertes.fr/hal-00872004

R. P. Kipling, P. Virkajärvi, L. Breitsameter, Y. Curnel, T. De-swaef et al., Key challenges and priorities for modelling European grasslands under climate change, Sci. Total Environ, vol.566, pp.851-864, 2016.
URL : https://hal.archives-ouvertes.fr/hal-01512163

J. Kattge, S. Díaz, S. Lavorel, I. C. Prentice, P. Leadley et al., TRY-A global database of plant traits, Glob. Chang. Biol, vol.17, pp.2905-2935, 2011.
URL : https://hal.archives-ouvertes.fr/hal-00639535

L. M. Verheijen, R. Aerts, V. Brovkin, J. Cavender-bares, J. H. Cornelissen et al., Inclusion of ecologically based trait variation in plant functional types reduces the projected land carbon sink in an earth system model, Glob. Chang. Biol, vol.21, pp.3074-3086, 2015.

M. Peaucelle, C. Bacour, P. Ciais, N. Vuichard, S. Kuppel et al., Covariations between plant functional traits emerge from constraining parameterization of a terrestrial biosphere model, Glob. Ecol. Biogeogr, vol.28, p.25, 2019.
URL : https://hal.archives-ouvertes.fr/hal-02371849

M. Van-oijen, Bayesian methods for quantifying and reducing uncertainty and error in forest models, Curr. For. Rep, vol.3, pp.269-280, 2017.

P. Cruz, J. Theau, E. Lecloux, C. Jouany, and M. Duru, Typologie fonctionnelle de graminées fourragères pérennes: Une classification multitraits, vol.201, pp.11-17, 2010.

V. Maire, P. Martre, J. Kattge, F. Gastal, G. Esser et al., The coordination of leaf photosynthesis links C and N Fluxes in C3 plant species, PLoS ONE, vol.7, 2012.
URL : https://hal.archives-ouvertes.fr/hal-01190237

I. C. Prentice, N. Dong, S. M. Gleason, V. Maire, and I. J. Wright, Balancing the costs of carbon gain and water transport: Testing a new theoretical framework for plant functional ecology, Ecol. Lett, vol.17, pp.82-91, 2014.

T. Chen, L. Cabrera-bosquet, S. Alvarez-prado, R. Perez, S. Artzet et al., Genetic and environmental dissection of biomass accumulation in multi-genotype maize canopies, J. Exp. Bot, vol.70, pp.2523-2534, 2019.
URL : https://hal.archives-ouvertes.fr/hal-01895279

M. Van-oijen, Tools for landscape science: Theory, models and data, Current Trends in Landscape Research

L. Müller and F. Eulenstein, , pp.219-231, 2019.

P. Levy, J. Drewer, M. Jammet, S. Leeson, T. Friborg et al., Inference of spatial heterogeneity in surface fluxes from eddy covariance data: A case study from a subarctic mire ecosystem, Agric. For. Meteorol, vol.280, 2020.

N. Puche, N. Senapati, C. R. Flechard, K. Klumpp, M. U. Kirschbaum et al., Modeling carbon and water fluxes of managed grasslands: Comparing flux variability and net carbon budgets between grazed and mowed systems, Agronomy, vol.9, p.183, 2019.
URL : https://hal.archives-ouvertes.fr/hal-02101575

J. Gurevitch, J. Koricheva, S. Nakagawa, and G. Stewart, Meta-analysis and the science of research synthesis, Nature, vol.555, pp.175-182, 2018.

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