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M. Oggenfuss, M. Holterman, and A. Both, Julie Cazareth from "Imaging and Cytometry platform" at IPMC for tech support. GDK has received the support of the EU in the framework of the Marie-Curie FP7 COFUND People Programme, through the award of an AgreenSkills+ fellowship (under grant number 609398). CMJ received funding from the ASEXEVOL project, grant number 392 ANR-13-JSV7-0006 which also funded bioinformatics equipment used for this paper. This work has been supported by the French government, through the UCA-JEDI "Investments in the Future" project managed by the National Research Agency (ANR) with the reference number ANR-15-IDEX-01, DNA extraction and quantification

E. G. , S. K. , D. K. , M. P. , L. P. B-;-l et al., performed measurements of the nuclear DNA content and analysed the results. G.D.K. cleaned and assembled the genome and participated in the analysis of the genome structure. E.G.J.D. participated in the analysis of the genome structure and performed the comparative assessment of genome completeness. M.P. performed the genome structure and divergence analysis. E.S. performed the gene prediction and analysed the results. D.K.K. performed the transposable elements annotation and analysis