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The Farm Animal Genotype–Tissue Expression (FarmGTEx) Project Lingzhao Fang a80064b573c78bf1ce89d8910635b223 igmm.ed.ac.uk 1255851 1169791-1255851 https://orcid.org/0000-0003-1103-3679 Jinyan Teng 1388800 3096080-1388800 https://orcid.org/0000-0002-3046-1452 Qing Lin 1064919-0 Zhonghao Bai 3165690-0 Shuli Liu 1437949-0 Dailu Guan 1388802 2009110-1388802 https://orcid.org/0000-0001-8800-3158 Bingjie Li c6b9224e9d5a211a03ec041d42b88295 sruc.ac.uk 1362067 1668456-1362067 https://orcid.org/0000-0002-8528-1889 Yahui Gao 1876430-0 Yali Hou 845214-0 Mian Gong 1178470 2057003-1178470 https://orcid.org/0000-0002-1785-0621 Zhangyuan Pan 3165694-0 Ying Yu 1255842 960189-1255842 https://orcid.org/0000-0002-4524-0791 Emily Clark 2109452-0 Jacqueline Smith 0ff72363b27a2eca0c7adeb0ded7559f roslin.ed.ac.uk 1287218 1871082-1287218 https://orcid.org/0000-0002-2813-7872 Konrad Rawlik 1255846 1100756-1255846 https://orcid.org/0000-0002-0010-370X Ruidong Xiang 1255843 2000222-1255843 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635124-1520627 https://orcid.org/0000-0002-6192-3170 Dan Hao 3448117-0 Lei Liu 772581-0 Siqian Chen 3448118-0 Robert Young 1520628 326280-1520628 https://orcid.org/0000-0002-8300-8002 Xia Shen 1237087 1043063-1237087 https://orcid.org/0000-0003-4390-1979 Charley Xia 1520629 2815534-1520629 https://orcid.org/0000-0002-8194-9351 Hao Cheng 1042925-0 Li Ma 1255848 721739-1255848 https://orcid.org/0000-0003-1038-1081 John Cole 1255849 958257-1255849 https://orcid.org/0000-0003-1242-4401 Ransom Baldwin 1520630 3448119-1520630 https://orcid.org/0000-0002-0753-4377 Cong-Jun Li 1255847 3165708-1255847 https://orcid.org/0000-0003-1389-9820 Curtis van Tassell 771878 844078-771878 https://orcid.org/0000-0002-8416-2087 Benjamin Rosen 19c53cb44515fc9b923ef37f5aa2d712 usda.gov 801139 1899400-801139 https://orcid.org/0000-0001-9395-8346 Nayan Bhowmik 1520631 3448120-1520631 https://orcid.org/0000-0003-3906-1460 Joan Lunney 1520632 939240-1520632 https://orcid.org/0000-0002-1147-8662 Wansheng Liu 1520633 745645-1520633 https://orcid.org/0000-0003-1788-7093 Leluo Guan 1434605-0 Xin Zhao 1520634 235928-1520634 https://orcid.org/0000-0002-1078-7332 Eveline Ibeagha-Awemu 1520635 764258-1520635 https://orcid.org/0000-0001-8810-9294 Yonglun Luo 1224615 2076529-1224615 https://orcid.org/0000-0002-0007-7759 Lin Lin 1388809 444970-1388809 https://orcid.org/0000-0002-7546-4948 Oriol Canela-Xandri 2815531-0 Martijn Derks 1788945-0 Richard Crooijmans 737514-0 Marta Gòdia 2106013-0 Ole Madsen 246770-0 Martien Groenen ea8e0440f62c96cd8d1c4178d4b0b8b9 wur.nl 946066 737518-946066 https://orcid.org/0000-0003-0484-4545 James Koltes 1347870 3141508-1347870 https://orcid.org/0000-0003-1897-5685 Christopher Tuggle 1388808 2104608-1388808 https://orcid.org/0000-0002-4229-5316 Fiona Mccarthy 1488917-0 Dominique Rocha 18546-0 Elisabetta Giuffra 002761509feba67e00f75fe2fb841366 inrae.fr elisabetta-giuffra 736095 49934-736095 http://www.researcherid.com/rid/J-8080-2014 https://orcid.org/0000-0001-9568-2056 https://scholar.google.it/citations?hl=fr&user=SpQe8sQAAAAJ http://www.researcherid.com/rid/http://www.researcherid.com/rid/J-8080-2014 Marcel Amills 769948 764571-769948 https://orcid.org/0000-0002-8999-0770 https://www.idref.fr/193655691 Alex Clop 1388807 764445-1388807 https://orcid.org/0000-0001-9238-2728 Maria Ballester e809ce6e062e25251ea4ae664022ce8b irta.cat 984898 777027-984898 https://orcid.org/0000-0002-5413-4640 Gwenola Tosser-Klopp 855b681ae3f5b821ea6c1de503ab228b inrae.fr gwenola-tosser 737163 9188-737163 https://orcid.org/0000-0003-0550-4673 https://www.idref.fr/067312497 Jing Li 243871-0 Chao Fang 636992-0 Ming Fang 1520636 721602-1520636 https://orcid.org/0000-0003-3066-4855 Qishan Wang 1388815 3096267-1388815 https://orcid.org/0000-0002-6475-0009 Zhuocheng Hou 1520637 3448121-1520637 https://orcid.org/0000-0003-4752-2255 Qin Wang 446689-0 Fuping Zhao 1520638 1038806-1520638 https://orcid.org/0000-0001-9582-5993 Lin Jiang 1178475 664155-1178475 https://orcid.org/0000-0001-9123-5988 Guiping Zhao 1520639 2146461-1520639 https://orcid.org/0000-0002-1510-6903 Zhengkui Zhou 1520640 3114880-1520640 https://orcid.org/0000-0002-5192-2465 Rong Zhou 1520641 1423801-1520641 https://orcid.org/0000-0001-6579-2641 Hehe Liu 3114876-0 Juan Deng 3448122-0 Long Jin 1520642 3448123-1520642 https://orcid.org/0000-0002-6055-4692 Mingzhou Li 1388810 3165712-1388810 https://orcid.org/0000-0001-8681-7684 Delin Mo 1520643 3448124-1520643 https://orcid.org/0000-0002-8738-4486 Xiaohong Liu 177239-0 Yaosheng Chen 3165716-0 Xiaolong Yuan 3165715-0 Jiaqi Li 1164779-0 Shuhong Zhao 1388813 745681-1388813 https://orcid.org/0000-0002-3997-2320 Yi Zhang 2532793-0 Xiangdong Ding 764809-0 Dongxiao Sun 1038811-0 Hui-Zeng Sun 1520644 3448125-1520644 https://orcid.org/0000-0001-5380-6030 Cong Li 995002-0 Yu Wang 2522858-0 Yu Jiang 1178471 2548326-1178471 https://orcid.org/0000-0003-4821-3585 Dongdong Wu 1178467 3389902-1178467 https://orcid.org/0000-0001-7101-7297 Wenwen Wang 950821-0 Xinzhong Fan 1199388-0 Qin Zhang 413255-0 Kui Li 1388816 764122-1388816 https://orcid.org/0000-0002-5686-3898 Hao Zhang 738131-0 Ning Yang d3e6895732a2d3f59abe8bab7ab4382e cau.edu.cn 984719 1042934-984719 https://orcid.org/0000-0001-5772-3320 Xiaoxiang Hu 297d7f52d7dabcc60d4ea70f7812df79 cau.edu.cn 956970 845141-956970 https://orcid.org/0000-0001-7045-0283 Wen Huang 1407607 1202268-1407607 https://orcid.org/0000-0001-6788-8364 Jiuzhou Song 845218-0 Yang Wu 304544-0 Jian Yang 1215658 203908-1215658 https://orcid.org/0000-0003-2001-2474 Weiwei Wu 3448126-0 Claudia Kasper c9aafb826ecae8f3ab755bee112cb450 agroscope.admin.ch 1367133 405309-1367133 https://orcid.org/0000-0001-7305-3996 Xinfeng Liu 1758233-0 Xiaofei Yu 3131638-0 Leilei Cui 1042755-0 Xiang Zhou 1274437 1276129-1274437 https://orcid.org/0000-0002-4331-7599 Seyoung Kim 3448127-0 Wei Li a9a21748c54da118354cdea1c0611a24 uci.edu 1390247 100222-1390247 https://orcid.org/0000-0001-9931-5990 Hae Kyung Im 1093149-0 Edward Buckler 1220274 1810851-1220274 https://orcid.org/0000-0002-3100-371X Bing Ren 1520645 1248918-1520645 https://orcid.org/0000-0002-5435-1127 Michael Schatz 1187271 212718-1187271 https://orcid.org/0000-0002-4118-4446 Jingyi Jessica Li 1520646 3448128-1520646 https://orcid.org/0000-0002-9288-5648 Abraham Palmer 1520647 1991397-1520647 https://orcid.org/0000-0003-3634-0747 Laurent Frantz 27eea5e35d5ddc8de6712f00c0959b49 gmail.com 771169 816151-771169 https://orcid.org/0000-0001-8030-3885 Huaijun Zhou 1388814 939421-1388814 https://orcid.org/0000-0001-6023-9521 Zhe Zhang c7d6aa3b4316154240a3d73761f80403 scau.edu.cn 1056994 1040606-1056994 https://orcid.org/0000-0001-7338-7718 George Liu 1388817 2528573-1388817 https://orcid.org/0000-0003-0192-6705 GWENOLA TOSSER-KLOPP 855b681ae3f5b821ea6c1de503ab228b inrae.fr 2025-03-17 12:38:08 2026-04-23 15:38:50 2025-03-17 12:38:08 2025-03-17 publisher version Scientists contributor GWENOLA TOSSER-KLOPP 855b681ae3f5b821ea6c1de503ab228b inrae.fr CCSD hal-04993821 https://hal.inrae.fr/hal-04993821 fang:hal-04993821 <i>Nature Genetics</i>, 2025, <a target="_blank" href="https://dx.doi.org/10.1038/s41588-025-02121-5">⟨10.1038/s41588-025-02121-5⟩</a> Nature Genetics, 2025, ⟨10.1038/s41588-025-02121-5⟩ AgroParisTech Ecole Nationale Supérieure Agronomique de Toulouse Université Paris-Saclay Archive ouverte en agrobiosciences Institut National de Recherche en Agriculture, Alimentation et Environnement Université Paris-Saclay Département GA INRAE Génétique Physiologie et Systèmes d'Elevage Graduate School Biologie, Société, Ecologie & Environnement, Ressources, Agriculture & Alimentation Graduate School Life Sciences and Health INRAE Occitanie Toulouse Toulouse - INP Université de Toulouse / Toulouse INP Université de Toulouse / Toulouse INP Génétique Animale et Biologie Intégrative Sciences Animales Paris-Saclay École Nationale Vétérinaire de Toulouse APT_TEST International No Yes The Farm Animal Genotype–Tissue Expression (FarmGTEx) Project Lingzhao Fang a80064b573c78bf1ce89d8910635b223 igmm.ed.ac.uk 1255851 1169791-1255851 https://orcid.org/0000-0003-1103-3679 Jinyan Teng 1388800 3096080-1388800 https://orcid.org/0000-0002-3046-1452 Qing Lin 1064919-0 Zhonghao Bai 3165690-0 Shuli Liu 1437949-0 Dailu Guan 1388802 2009110-1388802 https://orcid.org/0000-0001-8800-3158 Bingjie Li c6b9224e9d5a211a03ec041d42b88295 sruc.ac.uk 1362067 1668456-1362067 https://orcid.org/0000-0002-8528-1889 Yahui Gao 1876430-0 Yali Hou 845214-0 Mian Gong 1178470 2057003-1178470 https://orcid.org/0000-0002-1785-0621 Zhangyuan Pan 3165694-0 Ying Yu 1255842 960189-1255842 https://orcid.org/0000-0002-4524-0791 Emily Clark 2109452-0 Jacqueline Smith 0ff72363b27a2eca0c7adeb0ded7559f roslin.ed.ac.uk 1287218 1871082-1287218 https://orcid.org/0000-0002-2813-7872 Konrad Rawlik 1255846 1100756-1255846 https://orcid.org/0000-0002-0010-370X Ruidong Xiang 1255843 2000222-1255843 https://orcid.org/0000-0002-1584-7605 Amanda Chamberlain 1255844 844082-1255844 https://orcid.org/0000-0002-9395-1299 Michael Goddard 1186906-0 Mathew Littlejohn 2883613-0 Greger Larson 769740 745641-769740 https://orcid.org/0000-0002-4092-0392 David Machugh 1520624 1623564-1520624 https://orcid.org/0000-0002-8112-4704 John O’grady 2445307-0 Peter Sørensen 764542-0 Goutam Sahana 1263658 764669-1263658 https://orcid.org/0000-0001-7608-7577 Mogens Sandø Lund 1043098-0 Zhihua Jiang 1520625 1685332-1520625 https://orcid.org/0000-0003-1986-088X Xiangchun Pan 3165706-0 Wentao Gong 3448114-0 Haihan Zhang 3448115-0 Xi He 1008228-0 Yuebo Zhang 3448116-0 Ning Gao 1744503-0 Jun He 511267-0 Guoqiang Yi 1388811 1042928-1388811 https://orcid.org/0000-0001-8248-9126 Yuwen Liu 1520626 3096104-1520626 https://orcid.org/0009-0001-0470-2355 Zhonglin Tang 1388812 3096116-1388812 https://orcid.org/0000-0002-4538-4349 Pengju Zhao 3096330-0 Yang Zhou 397156-0 Liangliang Fu 803089-0 Xiao Wang 1520627 635124-1520627 https://orcid.org/0000-0002-6192-3170 Dan Hao 3448117-0 Lei Liu 772581-0 Siqian Chen 3448118-0 Robert Young 1520628 326280-1520628 https://orcid.org/0000-0002-8300-8002 Xia Shen 1237087 1043063-1237087 https://orcid.org/0000-0003-4390-1979 Charley Xia 1520629 2815534-1520629 https://orcid.org/0000-0002-8194-9351 Hao Cheng 1042925-0 Li Ma 1255848 721739-1255848 https://orcid.org/0000-0003-1038-1081 John Cole 1255849 958257-1255849 https://orcid.org/0000-0003-1242-4401 Ransom Baldwin 1520630 3448119-1520630 https://orcid.org/0000-0002-0753-4377 Cong-Jun Li 1255847 3165708-1255847 https://orcid.org/0000-0003-1389-9820 Curtis van Tassell 771878 844078-771878 https://orcid.org/0000-0002-8416-2087 Benjamin Rosen 19c53cb44515fc9b923ef37f5aa2d712 usda.gov 801139 1899400-801139 https://orcid.org/0000-0001-9395-8346 Nayan Bhowmik 1520631 3448120-1520631 https://orcid.org/0000-0003-3906-1460 Joan Lunney 1520632 939240-1520632 https://orcid.org/0000-0002-1147-8662 Wansheng Liu 1520633 745645-1520633 https://orcid.org/0000-0003-1788-7093 Leluo Guan 1434605-0 Xin Zhao 1520634 235928-1520634 https://orcid.org/0000-0002-1078-7332 Eveline Ibeagha-Awemu 1520635 764258-1520635 https://orcid.org/0000-0001-8810-9294 Yonglun Luo 1224615 2076529-1224615 https://orcid.org/0000-0002-0007-7759 Lin Lin 1388809 444970-1388809 https://orcid.org/0000-0002-7546-4948 Oriol Canela-Xandri 2815531-0 Martijn Derks 1788945-0 Richard Crooijmans 737514-0 Marta Gòdia 2106013-0 Ole Madsen 246770-0 Martien Groenen ea8e0440f62c96cd8d1c4178d4b0b8b9 wur.nl 946066 737518-946066 https://orcid.org/0000-0003-0484-4545 James Koltes 1347870 3141508-1347870 https://orcid.org/0000-0003-1897-5685 Christopher Tuggle 1388808 2104608-1388808 https://orcid.org/0000-0002-4229-5316 Fiona Mccarthy 1488917-0 Dominique Rocha 18546-0 Elisabetta Giuffra 002761509feba67e00f75fe2fb841366 inrae.fr elisabetta-giuffra 736095 49934-736095 http://www.researcherid.com/rid/J-8080-2014 https://orcid.org/0000-0001-9568-2056 https://scholar.google.it/citations?hl=fr&user=SpQe8sQAAAAJ http://www.researcherid.com/rid/http://www.researcherid.com/rid/J-8080-2014 Marcel Amills 769948 764571-769948 https://orcid.org/0000-0002-8999-0770 https://www.idref.fr/193655691 Alex Clop 1388807 764445-1388807 https://orcid.org/0000-0001-9238-2728 Maria Ballester e809ce6e062e25251ea4ae664022ce8b irta.cat 984898 777027-984898 https://orcid.org/0000-0002-5413-4640 Gwenola Tosser-Klopp 855b681ae3f5b821ea6c1de503ab228b inrae.fr gwenola-tosser 737163 9188-737163 https://orcid.org/0000-0003-0550-4673 https://www.idref.fr/067312497 Jing Li 243871-0 Chao Fang 636992-0 Ming Fang 1520636 721602-1520636 https://orcid.org/0000-0003-3066-4855 Qishan Wang 1388815 3096267-1388815 https://orcid.org/0000-0002-6475-0009 Zhuocheng Hou 1520637 3448121-1520637 https://orcid.org/0000-0003-4752-2255 Qin Wang 446689-0 Fuping Zhao 1520638 1038806-1520638 https://orcid.org/0000-0001-9582-5993 Lin Jiang 1178475 664155-1178475 https://orcid.org/0000-0001-9123-5988 Guiping Zhao 1520639 2146461-1520639 https://orcid.org/0000-0002-1510-6903 Zhengkui Zhou 1520640 3114880-1520640 https://orcid.org/0000-0002-5192-2465 Rong Zhou 1520641 1423801-1520641 https://orcid.org/0000-0001-6579-2641 Hehe Liu 3114876-0 Juan Deng 3448122-0 Long Jin 1520642 3448123-1520642 https://orcid.org/0000-0002-6055-4692 Mingzhou Li 1388810 3165712-1388810 https://orcid.org/0000-0001-8681-7684 Delin Mo 1520643 3448124-1520643 https://orcid.org/0000-0002-8738-4486 Xiaohong Liu 177239-0 Yaosheng Chen 3165716-0 Xiaolong Yuan 3165715-0 Jiaqi Li 1164779-0 Shuhong Zhao 1388813 745681-1388813 https://orcid.org/0000-0002-3997-2320 Yi Zhang 2532793-0 Xiangdong Ding 764809-0 Dongxiao Sun 1038811-0 Hui-Zeng Sun 1520644 3448125-1520644 https://orcid.org/0000-0001-5380-6030 Cong Li 995002-0 Yu Wang 2522858-0 Yu Jiang 1178471 2548326-1178471 https://orcid.org/0000-0003-4821-3585 Dongdong Wu 1178467 3389902-1178467 https://orcid.org/0000-0001-7101-7297 Wenwen Wang 950821-0 Xinzhong Fan 1199388-0 Qin Zhang 413255-0 Kui Li 1388816 764122-1388816 https://orcid.org/0000-0002-5686-3898 Hao Zhang 738131-0 Ning Yang d3e6895732a2d3f59abe8bab7ab4382e cau.edu.cn 984719 1042934-984719 https://orcid.org/0000-0001-5772-3320 Xiaoxiang Hu 297d7f52d7dabcc60d4ea70f7812df79 cau.edu.cn 956970 845141-956970 https://orcid.org/0000-0001-7045-0283 Wen Huang 1407607 1202268-1407607 https://orcid.org/0000-0001-6788-8364 Jiuzhou Song 845218-0 Yang Wu 304544-0 Jian Yang 1215658 203908-1215658 https://orcid.org/0000-0003-2001-2474 Weiwei Wu 3448126-0 Claudia Kasper c9aafb826ecae8f3ab755bee112cb450 agroscope.admin.ch 1367133 405309-1367133 https://orcid.org/0000-0001-7305-3996 Xinfeng Liu 1758233-0 Xiaofei Yu 3131638-0 Leilei Cui 1042755-0 Xiang Zhou 1274437 1276129-1274437 https://orcid.org/0000-0002-4331-7599 Seyoung Kim 3448127-0 Wei Li a9a21748c54da118354cdea1c0611a24 uci.edu 1390247 100222-1390247 https://orcid.org/0000-0001-9931-5990 Hae Kyung Im 1093149-0 Edward Buckler 1220274 1810851-1220274 https://orcid.org/0000-0002-3100-371X Bing Ren 1520645 1248918-1520645 https://orcid.org/0000-0002-5435-1127 Michael Schatz 1187271 212718-1187271 https://orcid.org/0000-0002-4118-4446 Jingyi Jessica Li 1520646 3448128-1520646 https://orcid.org/0000-0002-9288-5648 Abraham Palmer 1520647 1991397-1520647 https://orcid.org/0000-0003-3634-0747 Laurent Frantz 27eea5e35d5ddc8de6712f00c0959b49 gmail.com 771169 816151-771169 https://orcid.org/0000-0001-8030-3885 Huaijun Zhou 1388814 939421-1388814 https://orcid.org/0000-0001-6023-9521 Zhe Zhang c7d6aa3b4316154240a3d73761f80403 scau.edu.cn 1056994 1040606-1056994 https://orcid.org/0000-0001-7338-7718 George Liu 1388817 2528573-1388817 https://orcid.org/0000-0003-0192-6705 7336 1061-4036 1546-1718 Nature Genetics Nature Publishing Group 2025-03-17 10.1038/s41588-025-02121-5 001453159700001 English Life Sciences [q-bio] Life Sciences [q-bio]/Biochemistry, Molecular Biology/Genomics [q-bio.GN] Journal articles Journal articles Journal articles The Roslin Institute

Easter Bush Midlothian EH25 9RG Scotland, UK

http://www.roslin.ed.ac.uk/ USDA Agricultural Research Service [Beltsville, Maryland]

Beltsville Agricultural Research Center10300 Baltimore AvenueBldg. 003, Rm. 231, BARC-WestBeltsville, MD 20705

https://www.ars.usda.gov/northeast-area/beltsville-md-barc/beltsville-agricultural-research-center/ 200917790C https://ror.org/03rkgeb39 Génétique Animale et Biologie Intégrative GABI 2020-01-01

Domaine de Vilvert F-78252 Jouy-en-Josas

https://gabi.jouy.hub.inrae.fr/ Departament de Matemàtiques [Barcelona] UAB

Universitat Autònoma de Barcelona, Edifici C Facultat de Ciències, 08193 Bellaterra - Barcelona

https://www.uab.cat/matematiques/ 201421780T Génétique Physiologie et Systèmes d'Elevage GenPhySE 2025-01-01

INRAE Auzeville - 24, chemin de Borde-Rouge -Auzeville Tolosane31326 Castanet Tolosan

http://genphyse.toulouse.inra.fr/ Biotechnology and Biological Sciences Research Council BBSRC

https://www.ukri.org/councils/bbsrc/ https://ror.org/02d2m2044 USDA-ARS : Agricultural Research Service

https://www.ars.usda.gov/ 139408088 https://ror.org/02kbmgc12 AgroParisTech 2007-01-01

22 place de l'Agronomie CS 20040 91123 Palaiseau cedex

https://www.agroparistech.fr/ 241345251 https://ror.org/03xjwb503 Université Paris-Saclay

Bâtiment Bréguet, 3 Rue Joliot Curie 2e ét, 91190 Gif-sur-Yvette

https://www.universite-paris-saclay.fr/fr https://ror.org/003vg9w96 Institut National de Recherche pour l’Agriculture, l’Alimentation et l’Environnement INRAE 2020-01-01

0000 0001 2296 0625 https://ror.org/052g8jq94 Universitat Autònoma de Barcelona = Autonomous University of Barcelona = Universidad Autónoma de Barcelona UAB 1968-01-01

UAB Campus 08193 Bellaterra Barcelona

https://www.uab.cat/ https://ror.org/03m3gzv89 École nationale vétérinaire de Toulouse ENVT

23 chemin des Capelles - 31076 Toulouse Cedex

https://envt.fr/ 026388820 0000000121766169 https://ror.org/033p9g875 Institut National Polytechnique (Toulouse) Toulouse INP

https://www.inp-toulouse.fr/ https://ror.org/017tgbk05 Communauté d'universités et établissements de Toulouse Comue de Toulouse

41 Allée Jules Guesde, 31000 Toulouse

https://www.univ-toulouse.fr/ 0000 0001 2182 8729 https://ror.org/03d5w9863 École nationale supérieure agronomique de Toulouse ENSAT, INP-ENSAT, AgroToulouse, INP-AgroToulouse

https://www.ensat.fr/fr/index.html Ecole d'Ingénieurs de Purpan EI Purpan 2025-01-01

75, voie du TOEC - BP 5761131076 Toulouse

https://www.purpan.fr/