J. Parkinson, Signal transduction schemes of bacteria, Cell, vol.73, issue.5, pp.857-871, 1993.
DOI : 10.1016/0092-8674(93)90267-T

T. Oshima, H. Aiba, Y. Masuda, S. Kanaya, and M. Sugiura, Transcriptome analysis of all two-component regulatory system mutants of Escherichia coli K-12, Molecular Microbiology, vol.97, issue.1, pp.281-291, 2002.
DOI : 10.1046/j.1365-2958.2002.03170.x

B. F. Wanner, R. Neidhardt, J. L. Ingraham, E. C. Lin, K. B. Low et al., Phosphorus assimilation and control of the phosphate regulon, Escherichia coli and Salmonella: Cellular and Molecular Biology, pp.1357-1381, 1996.

N. Rao and A. Torriani, Molecular aspects of phosphate transport in Escherichia coli, Molecular Microbiology, vol.168, issue.7, pp.1083-1090, 1990.
DOI : 10.1007/BF00391740

F. Chan and A. Torriani, PstB protein of the phosphate-specific transport system of Escherichia coli is an ATPase., Journal of Bacteriology, vol.178, issue.13, pp.3974-3977, 1996.
DOI : 10.1128/jb.178.13.3974-3977.1996

Y. Hsieh and B. Wanner, Global regulation by the seven-component Pi signaling system, Current Opinion in Microbiology, vol.13, issue.2, pp.198-203, 2010.
DOI : 10.1016/j.mib.2010.01.014

C. Yang, T. Huang, S. Wen, C. Chang, and S. Tsai, Genome-Wide PhoB Binding and Gene Expression Profiles Reveal the Hierarchical Gene Regulatory Network of Phosphate Starvation in Escherichia coli, PLoS ONE, vol.7, issue.10, p.47314, 2012.
DOI : 10.1371/journal.pone.0047314.s008

Y. Yoshida, S. Sugiyama, T. Oyamada, K. Yokoyama, and K. Makino, Novel members of the phosphate regulon in Escherichia coli O157:H7 identified using a whole-genome shotgun approach, Gene, vol.502, issue.1, pp.27-35, 2012.
DOI : 10.1016/j.gene.2012.03.064

B. Wanner, Molecular genetics of carbon-phosphorus bond cleavage in bacteria, Biodegradation, vol.58, issue.3-4, pp.175-184, 1994.
DOI : 10.1007/BF00696458

C. Cheng, M. Wakefield, J. Yang, M. Tauschek, and R. Robins-browne, Genome-Wide Analysis of the Pho Regulon in a pstCA Mutant of Citrobacter rodentium, PLoS ONE, vol.174, issue.11, p.50682, 2012.
DOI : 10.1371/journal.pone.0050682.s006

S. Crepin, M. Lamarche, P. Garneau, J. Seguin, and J. Proulx, Genome-wide transcriptional response of an avian pathogenic Escherichia coli (APEC) pst mutant, BMC Genomics, vol.9, issue.1, p.568, 2008.
DOI : 10.1186/1471-2164-9-568
URL : https://hal.archives-ouvertes.fr/pasteur-00720793

T. Danhorn, M. Hentzer, M. Givskov, M. Parsek, and C. Fuqua, Phosphorus Limitation Enhances Biofilm Formation of the Plant Pathogen Agrobacterium tumefaciens through the PhoR-PhoB Regulatory System, Journal of Bacteriology, vol.186, issue.14, pp.4492-4501, 2004.
DOI : 10.1128/JB.186.14.4492-4501.2004

M. Lamarche, C. Dozois, F. Daigle, M. Caza, and R. Curtiss, Inactivation of the Pst System Reduces the Virulence of an Avian Pathogenic Escherichia coli O78 Strain, Infection and Immunity, vol.73, issue.7, pp.4138-4145, 2005.
DOI : 10.1128/IAI.73.7.4138-4145.2005
URL : https://hal.archives-ouvertes.fr/pasteur-00720810

O. Zaborina, C. Holbrook, Y. Chen, J. Long, and A. Zaborin, Structure???Function Aspects of PstS in Multi-Drug???Resistant Pseudomonas aeruginosa, PLoS Pathogens, vol.148, issue.2, p.43, 2008.
DOI : 10.1371/journal.ppat.0040043.sg004

N. Bertrand, S. Houle, G. Lebihan, E. Poirier, and C. Dozois, Increased Pho Regulon Activation Correlates with Decreased Virulence of an Avian Pathogenic Escherichia coli O78 Strain, Infection and Immunity, vol.78, issue.12, pp.5324-5331, 2010.
DOI : 10.1128/IAI.00452-10
URL : https://hal.archives-ouvertes.fr/pasteur-00720787

S. Crepin, S. Chekabab, L. Bihan, G. Bertrand, N. Dozois et al., The Pho regulon and the pathogenesis of Escherichia coli, Veterinary Microbiology, vol.153, issue.1-2, pp.82-88, 2011.
DOI : 10.1016/j.vetmic.2011.05.043
URL : https://hal.archives-ouvertes.fr/pasteur-00720774

P. Griffin and R. Tauxe, The Epidemiology of Infections Caused by Escherichia coli O157: H7, Other Enterohemorrhagic E. coli, and the Associated Hemolytic Uremic Syndrome, Epidemiologic Reviews, vol.13, issue.1, pp.60-98, 1991.
DOI : 10.1093/oxfordjournals.epirev.a036079

C. Gyles, Shiga toxin-producing An overview, Journal of Animal Science, vol.85, issue.13_suppl, pp.45-62, 2007.
DOI : 10.2527/jas.2006-508

T. Mcdaniel, K. Jarvis, M. Donnenberg, and J. Kaper, A genetic locus of enterocyte effacement conserved among diverse enterobacterial pathogens., Proceedings of the National Academy of Sciences, vol.92, issue.5, pp.1664-1668, 1995.
DOI : 10.1073/pnas.92.5.1664

S. Elliott, V. Sperandio, J. Giron, S. Shin, and J. Mellies, The Locus of Enterocyte Effacement (LEE)-Encoded Regulator Controls Expression of Both LEE- and Non-LEE-Encoded Virulence Factors in Enteropathogenic and Enterohemorrhagic Escherichia coli, Infection and Immunity, vol.68, issue.11, pp.6115-6126, 2000.
DOI : 10.1128/IAI.68.11.6115-6126.2000

K. Haack, C. Robinson, K. Miller, J. Fowlkes, and J. Mellies, Interaction of Ler at the LEE5 (tir) Operon of Enteropathogenic Escherichia coli, Infection and Immunity, vol.71, issue.1, pp.384-392, 2003.
DOI : 10.1128/IAI.71.1.384-392.2003

V. Sperandio, J. Mellies, R. Delahay, G. Frankel, and J. Crawford, Activation of enteropathogenic Escherichia coli (EPEC) LEE2 and LEE3 operons by Ler, Molecular Microbiology, vol.49, issue.4, pp.781-793, 2000.
DOI : 10.1046/j.1365-2958.1998.00771.x

W. Deng, J. Puente, S. Gruenheid, Y. Li, and B. Vallance, Dissecting virulence: Systematic and functional analyses of a pathogenicity island, Proceedings of the National Academy of Sciences, vol.101, issue.10, pp.3597-3602, 2004.
DOI : 10.1073/pnas.0400326101

J. Tree, E. Wolfson, D. Wang, A. Roe, and D. Gally, Controlling injection: regulation of type III secretion in enterohaemorrhagic Escherichia coli, Trends in Microbiology, vol.17, issue.8, pp.361-370, 2009.
DOI : 10.1016/j.tim.2009.06.001

S. Bhatt, T. Romeo, and D. Kalman, Honing the message: post-transcriptional and post-translational control in attaching and effacing pathogens, Trends in Microbiology, vol.19, issue.5, pp.217-224, 2011.
DOI : 10.1016/j.tim.2011.01.004

J. Little, The SOS regulatory system Regulation of gene expression in Escherichia coli, pp.453-479, 1995.

D. Hancock, T. Besser, J. Lejeune, M. Davis, and D. Rice, The control of VTEC in the animal reservoir, International Journal of Food Microbiology, vol.66, issue.1-2, pp.71-78, 2001.
DOI : 10.1016/S0168-1605(00)00487-6

S. Chekabab, J. Paquin-veillette, C. Dozois, and J. Harel, O157:H7, FEMS Microbiology Letters, vol.341, issue.1, pp.1-12, 2013.
DOI : 10.1111/1574-6968.12078
URL : https://hal.archives-ouvertes.fr/pasteur-01130833

S. Chekabab, F. Daigle, S. Charette, C. Dozois, and J. Harel, Survival of enterohemorrhagic Escherichia coli in the presence of Acanthamoeba castellanii and its dependence on Pho regulon, pp.427-437, 2012.

M. Carruthers, B. Bellaire, and F. Minion, Exploring the response of Escherichia coli???O157:H7 EDL933 within Acanthamoeba castellanii???by genome-wide transcriptional profiling, FEMS Microbiology Letters, vol.312, issue.1, pp.15-23, 2010.
DOI : 10.1111/j.1574-6968.2010.02098.x

M. Lamarche, B. Wanner, S. Crepin, and J. Harel, The phosphate regulon and bacterial virulence: a regulatory network connecting phosphate homeostasis and pathogenesis, FEMS Microbiology Reviews, vol.32, issue.3, pp.461-473, 2008.
DOI : 10.1111/j.1574-6976.2008.00101.x

J. Pratt, A. Ismail, and A. Camilli, PhoB regulates both environmental and virulence gene expression in Vibrio cholerae, Molecular Microbiology, vol.17, issue.6, pp.1595-1605, 2010.
DOI : 10.1111/j.1365-2958.2010.07310.x

J. Magid, H. Tiessen, and L. Condron, Dynamics of Organic Phosphorus in Soils under Natural and Agricultural Ecosystems, Humic substances in terrestrial ecosystems, pp.429-466, 1996.
DOI : 10.1016/B978-044481516-3/50012-8

M. Zubkov, I. Mary, E. Woodward, P. Warwick, and B. Fuchs, Microbial control of phosphate in the nutrient-depleted North Atlantic subtropical gyre, Environmental Microbiology, vol.40, issue.8, pp.2079-2089, 2007.
DOI : 10.1111/j.1462-2920.2007.01246.x
URL : https://hal.archives-ouvertes.fr/hal-00528815

L. Kayne, D. Argenio, D. Meyer, J. Hu, M. Jamgotchian et al., Analysis of segmental phosphate absorption in intact rats. A compartmental analysis approach., Journal of Clinical Investigation, vol.91, issue.3, pp.915-922, 1993.
DOI : 10.1172/JCI116313

J. Walton and T. Gray, Absorption of Inorganic Phosphate in the Human Small Intestine, Clinical Science, vol.56, issue.5, pp.407-412, 1979.
DOI : 10.1042/cs0560407

J. Lemann, Intestinal absorption of calcium, magnesium, and phosphorus Primer on the metabolic bone diseases and disorders of mineral metabolism, pp.46-50, 1993.

R. Pieper, Q. Zhang, D. Clark, P. Parmar, and H. Alami, Proteomic View of Interactions of Shiga Toxin-Producing Escherichia coli with the Intestinal Environment in Gnotobiotic Piglets, PLoS ONE, vol.58, issue.6, p.66462, 2013.
DOI : 10.1371/journal.pone.0066462.s004

F. Daigle, J. Fairbrother, and J. Harel, Identification of a mutation in the pstphoU operon that reduces pathogenicity of an Escherichia coli strain causing septicemia in pigs, Infect Immun, vol.63, pp.4924-4927, 1995.

M. Lamarche and J. Harel, Membrane Homeostasis Requires Intact pst in Extraintestinal Pathogenic Escherichia coli, Current Microbiology, vol.6, issue.5, pp.356-359, 2010.
DOI : 10.1007/s00284-009-9549-x

S. Crepin, S. Houle, M. Charbonneau, M. Mourez, and J. Harel, Mutant of Uropathogenic Escherichia coli Reduces Urinary Tract Infection, Infection and Immunity, vol.80, issue.8, pp.2802-2815, 2012.
DOI : 10.1128/IAI.00162-12
URL : https://hal.archives-ouvertes.fr/pasteur-00719756

C. Cheng, S. Tennant, K. Azzopardi, V. Bennett-wood, and E. Hartland, Contribution of the pst-phoU Operon to Cell Adherence by Atypical Enteropathogenic Escherichia coli and Virulence of Citrobacter rodentium, Infection and Immunity, vol.77, issue.5, pp.1936-1944, 2009.
DOI : 10.1128/IAI.01246-08

M. Lamarche, S. Kim, S. Crepin, M. Mourez, and N. Bertrand, Modulation of Hexa-Acyl Pyrophosphate Lipid A Population under Escherichia coli Phosphate (Pho) Regulon Activation, Journal of Bacteriology, vol.190, issue.15, pp.5256-5264, 2008.
DOI : 10.1128/JB.01536-07

R. Graveline, E ´ tude des mécanismes moléculaires influençant la variation de phase des adhésines P, F1651 et CS31A présentes chez des souches d'Escherichia coli pathogènes. Faculty of veterinary medecine: Montreal, 2011.

I. Muhldorfer, J. Hacker, G. Keusch, D. Acheson, and H. Tschape, Regulation of the Shiga-like toxin II operon in Escherichia coli, Infect Immun, vol.64, pp.495-502, 1996.

T. De-sablet, Y. Bertin, M. Vareille, J. Girardeau, and A. Garrivier, Differential expression of stx2 variants in Shiga toxin-producing Escherichia coli belonging to seropathotypes A and C, Microbiology, vol.154, issue.1, pp.176-186, 2008.
DOI : 10.1099/mic.0.2007/009704-0

F. Neidhardt, P. Bloch, and D. Smith, Culture medium for enterobacteria, J Bacteriol, vol.119, pp.736-747, 1974.

J. Peterson, L. Umayam, T. Dickinson, E. Hickey, and O. White, The Comprehensive Microbial Resource, Nucleic Acids Research, vol.29, issue.1, pp.123-125, 2001.
DOI : 10.1093/nar/29.1.123

R. Arribas-bosacoma, S. Kim, C. Ferrer-orta, A. Blanco, and P. Pereira, The X-ray Crystal Structures of Two Constitutively Active Mutants of the Escherichia coli PhoB Receiver Domain Give Insights into Activation, Journal of Molecular Biology, vol.366, issue.2, pp.626-641, 2007.
DOI : 10.1016/j.jmb.2006.11.038

R. Graveline, M. Mourez, M. Hancock, C. Martin, and S. Boisclair, Lrp-DNA complex stability determines the level of ON cells in type P fimbriae phase variation, Molecular Microbiology, vol.30, issue.4, pp.1286-1299, 2011.
DOI : 10.1111/j.1365-2958.2011.07761.x

W. Deng, Y. Li, P. Hardwidge, E. Frey, and R. Pfuetzner, Regulation of Type III Secretion Hierarchy of Translocators and Effectors in Attaching and Effacing Bacterial Pathogens, Infection and Immunity, vol.73, issue.4, pp.2135-2146, 2005.
DOI : 10.1128/IAI.73.4.2135-2146.2005

D. Dasanayake, M. Richaud, N. Cyr, C. Caballero-franco, and S. Pittroff, The N-terminal amphipathic region of the Escherichia coli type III secretion system protein EspD is required for membrane insertion and function, Molecular Microbiology, vol.28, issue.3, pp.734-750, 2011.
DOI : 10.1111/j.1365-2958.2011.07727.x

A. Gobert, M. Vareille, A. Glasser, T. Hindre, and T. De-sablet, Shiga Toxin Produced by Enterohemorrhagic Escherichia coli Inhibits PI3K/NF-??B Signaling Pathway in Globotriaosylceramide-3-Negative Human Intestinal Epithelial Cells, The Journal of Immunology, vol.178, issue.12, pp.8168-8174, 2007.
DOI : 10.4049/jimmunol.178.12.8168

W. Deng, J. Puente, S. Gruenheid, Y. Li, and B. Vallance, Dissecting virulence: Systematic and functional analyses of a pathogenicity island, Proceedings of the National Academy of Sciences, vol.101, issue.10, pp.3597-3602, 2004.
DOI : 10.1073/pnas.0400326101

R. Hengge-aronis, Signal Transduction and Regulatory Mechanisms Involved in Control of the ??S (RpoS) Subunit of RNA Polymerase, Microbiology and Molecular Biology Reviews, vol.66, issue.3, pp.373-395, 2002.
DOI : 10.1128/MMBR.66.3.373-395.2002

P. Loewen, B. Hu, J. Strutinsky, and R. Sparling, Regulation in the <i>rpoS</i> regulon of <i>Escherichia coli</i>, Canadian Journal of Microbiology, vol.44, issue.8, pp.707-717, 1998.
DOI : 10.1139/cjm-44-8-707

A. Bougdour, S. Wickner, and S. Gottesman, Modulating RssB activity: IraP, a novel regulator of ??S stability in Escherichia coli, Genes & Development, vol.20, issue.7, pp.884-897, 2006.
DOI : 10.1101/gad.1400306

S. Altuvia, D. Weinstein-fischer, A. Zhang, L. Postow, and G. Storz, A Small, Stable RNA Induced by Oxidative Stress: Role as a Pleiotropic Regulator and Antimutator, Cell, vol.90, issue.1, pp.43-53, 1997.
DOI : 10.1016/S0092-8674(00)80312-8
URL : http://doi.org/10.1016/s0092-8674(00)80312-8

M. Mandel and T. Silhavy, Starvation for Different Nutrients in Escherichia coli Results in Differential Modulation of RpoS Levels and Stability, Journal of Bacteriology, vol.187, issue.2, pp.434-442, 2005.
DOI : 10.1128/JB.187.2.434-442.2005

S. Herold, H. Karch, and H. Schmidt, Shiga toxin-encoding bacteriophages ??? genomes in motion, International Journal of Medical Microbiology, vol.294, issue.2-3, pp.115-121, 2004.
DOI : 10.1016/j.ijmm.2004.06.023

A. Unkmeir and H. Schmidt, Structural Analysis of Phage-Borne stx Genes and Their Flanking Sequences in Shiga Toxin-Producing Escherichia coli and Shigella dysenteriae Type 1 Strains, Infection and Immunity, vol.68, issue.9, pp.4856-4864, 2000.
DOI : 10.1128/IAI.68.9.4856-4864.2000

J. Tyler, M. Mills, and D. Friedman, The Operator and Early Promoter Region of the Shiga Toxin Type 2-Encoding Bacteriophage 933W and Control of Toxin Expression, Journal of Bacteriology, vol.186, issue.22, pp.7670-7679, 2004.
DOI : 10.1128/JB.186.22.7670-7679.2004

A. Blanco, M. Sola, F. Gomis-ruth, and M. Coll, Tandem DNA Recognition by PhoB, a Two-Component Signal Transduction Transcriptional Activator, Structure, vol.10, issue.5, pp.701-713, 2002.
DOI : 10.1016/S0969-2126(02)00761-X

M. Kendall, D. Rasko, and V. Sperandio, The LysR-type regulator QseA regulates both characterized and putative virulence genes in enterohaemorrhagic Escherichia coli O157:H7, Molecular Microbiology, vol.72, issue.5, pp.1306-1321, 2010.
DOI : 10.1111/j.1365-2958.2010.07174.x

J. Mellies, A. Barron, and A. Carmona, Enteropathogenic and Enterohemorrhagic Escherichia coli Virulence Gene Regulation, Infection and Immunity, vol.75, issue.9, pp.4199-4210, 2007.
DOI : 10.1128/IAI.01927-06
URL : http://www.ncbi.nlm.nih.gov/pmc/articles/PMC1951183

R. Gao and A. Stock, Evolutionary Tuning of Protein Expression Levels of a Positively Autoregulated Two-Component System, PLoS Genetics, vol.423, issue.10, p.1003927, 2013.
DOI : 10.1371/journal.pgen.1003927.s006

R. Gao and A. Stock, Probing kinase and phosphatase activities of two-component systems in vivo with concentration-dependent phosphorylation profiling, Proceedings of the National Academy of Sciences, vol.110, issue.2, pp.672-677, 2013.
DOI : 10.1073/pnas.1214587110

Y. Yoshida, S. Sugiyama, T. Oyamada, K. Yokoyama, and K. Makino, Identification and characterization of novel phosphate regulon genes, Escherichia coli O157:H7, pp.540-0544, 2010.

S. Fisher, W. Jiang, B. Wanner, and C. Walsh, Cross-talk between the Histidine Protein Kinase VanS and the Response Regulator PhoB: CHARACTERIZATION AND IDENTIFICATION OF A VanS DOMAIN THAT INHIBITS ACTIVATION OF PhoB, Journal of Biological Chemistry, vol.270, issue.39, pp.23143-23149, 1995.
DOI : 10.1074/jbc.270.39.23143

E. Suziedeliene, K. Suziedelis, V. Garbenciute, and S. Normark, The acidinducible asr gene in Escherichia coli: transcriptional control by the phoBR operon, J Bacteriol, vol.181, pp.2084-2093, 1999.

B. Wanner and M. Wilmes-riesenberg, Involvement of phosphotransacetylase, acetate kinase, and acetyl phosphate synthesis in control of the phosphate regulon in Escherichia coli., Journal of Bacteriology, vol.174, issue.7, pp.2124-2130, 1992.
DOI : 10.1128/jb.174.7.2124-2130.1992

M. Waldor and D. Friedman, Phage regulatory circuits and virulence gene expression, Current Opinion in Microbiology, vol.8, issue.4, pp.459-465, 2005.
DOI : 10.1016/j.mib.2005.06.001

P. Wagner, M. Neely, X. Zhang, D. Acheson, and M. Waldor, Role for a Phage Promoter in Shiga Toxin 2 Expression from a Pathogenic Escherichia coli Strain, Journal of Bacteriology, vol.183, issue.6, pp.2081-2085, 2001.
DOI : 10.1128/JB.183.6.2081-2085.2001

D. Rozanov, D. Ari, R. Sineoky, and S. , RecA-independent pathways of lambdoid prophage induction in Escherichia coli, J Bacteriol, vol.180, pp.6306-6315, 1998.

P. Shkilnyj and G. Koudelka, Effect of Salt Shock on Stability of ??imm434 Lysogens, Journal of Bacteriology, vol.189, issue.8, pp.3115-3123, 2007.
DOI : 10.1128/JB.01857-06

D. Ghosh, K. Roy, K. Williamson, S. Srinivasiah, and K. Wommack, Acyl-Homoserine Lactones Can Induce Virus Production in Lysogenic Bacteria: an Alternative Paradigm for Prophage Induction, Applied and Environmental Microbiology, vol.75, issue.22, pp.7142-7152, 2009.
DOI : 10.1128/AEM.00950-09

L. Imamovic and M. Muniesa, Characterizing RecA-Independent Induction of Shiga toxin2-Encoding Phages by EDTA Treatment, PLoS ONE, vol.25, issue.2, p.32393, 2012.
DOI : 10.1371/journal.pone.0032393.s001

N. Strockbine, L. Marques, J. Newland, H. Smith, and R. Holmes, Two toxin-converting phages from Escherichia coli O157:H7 strain 933 encode antigenically distinct toxins with similar biologic activities, Infection and immunity, vol.53, pp.135-140, 1986.

H. Kang, J. Srinivasan, and R. Curtiss, Immune Responses to Recombinant Pneumococcal PspA Antigen Delivered by Live Attenuated Salmonella enterica Serovar Typhimurium Vaccine, Infection and Immunity, vol.70, issue.4, pp.1739-1749, 2002.
DOI : 10.1128/IAI.70.4.1739-1749.2002

T. Hashimoto-gotoh, F. Franklin, A. Nordheim, and K. Timmis, Specific-purpose plasmid cloning vectors I. Low copy number, temperature-sensitive, mobilization-defective pSC101-derived containment vectors, Gene, vol.16, issue.1-3, pp.227-235, 1981.
DOI : 10.1016/0378-1119(81)90079-2