C. Ammer, L. seedlings in response to shading and small changes in the R/FR-ratio of radiation, Annals of Forest Science, vol.60, issue.2, pp.163-171, 2003.
DOI : 10.1051/forest:2003009
URL : https://hal.archives-ouvertes.fr/hal-01269417

P. J. Aphalo, C. L. Ballare, and A. L. Scopel, Plant-plant signalling, the shade-avoidance response and competition, Journal of Experimental Botany, vol.50, issue.340, pp.1629-1634, 1999.
DOI : 10.1093/jxb/50.340.1629

G. Aussenac, Interactions between forest stands and microclimate: Ecophysiological aspects and consequences for silviculture, Annals of Forest Science, vol.57, issue.3, pp.287-301, 2000.
DOI : 10.1051/forest:2000119
URL : https://hal.archives-ouvertes.fr/hal-00883349

P. Balandier, G. Sonohat, H. Sinoquet, C. Varlet-grancher, and Y. Dumas, Characterisation, prediction and relationships between different wavebands of solar radiation transmitted in the understorey of even-aged oak (Quercus petraea, Q. robur) stands, Trees, vol.121, issue.3, pp.363-370, 2006.
DOI : 10.1007/s00468-006-0049-3
URL : https://hal.archives-ouvertes.fr/hal-01189187

P. Balandier, H. Sinoquet, E. Frak, R. Giuliani, M. Vandame et al., Six-year time course of light-use efficiency, carbon gain and growth of beech saplings (Fagus sylvatica) planted under a Scots pine (Pinus sylvestris) shelterwood, Tree Physiology, vol.27, issue.8, pp.1073-1082, 2007.
DOI : 10.1093/treephys/27.8.1073
URL : https://hal.archives-ouvertes.fr/hal-01189176

P. Balandier, A. Marquier, S. Perret, C. Collet, and B. Courbaud, Comment estimer la lumière dans le sous-bois forestier à partir des caractéristiques dendrométriques des peuplements ? Rendezvous Techniques ONF, pp.27-28, 2010.

C. L. Ballaré, Keeping up with the neighbours: phytochrome sensing and other signalling mechanisms, Trends in Plant Science, vol.4, issue.3, pp.97-102, 1999.
DOI : 10.1016/S1360-1385(99)01383-7

J. Bengtsson, S. G. Nilsson, A. Franc, and P. Menozzi, Biodiversity, disturbances, ecosystem function and management of European forests, Forest Ecology and Management, vol.132, issue.1, pp.39-50, 2000.
DOI : 10.1016/S0378-1127(00)00378-9

Y. Claveau, C. Messier, and P. G. Comeau, Interacting influence of light and size on aboveground biomass distribution in sub-boreal conifer saplings with contrasting shade tolerance, Tree Physiology, vol.25, issue.3, pp.373-384, 2005.
DOI : 10.1093/treephys/25.3.373

Y. Claveau, C. Messier, P. G. Comeau, and K. D. Coates, Growth and crown morphological responses of boreal conifer seedlings and saplings with contrasting shade tolerance to a gradient of light and height, Canadian Journal of Forest Research, vol.32, issue.3, pp.458-468, 2002.
DOI : 10.1139/x01-220

T. Curt, L. Coll, B. Prevosto, P. Balandier, and G. Kunstler, Plasticity in growth, biomass allocation and root morphology in beech seedlings as induced by irradiance and herbaceous competition, Annals of Forest Science, vol.62, issue.1, pp.51-60, 2005.
DOI : 10.1051/forest:2004092
URL : https://hal.archives-ouvertes.fr/hal-00883860

D. Chantal, M. Leinonen, K. Kuuluvainen, T. Cescatti, and A. , Early response of Pinus sylvestris and Picea abies seedlings to an experimental canopy gap in a boreal spruce forest, Forest Ecology and Management, vol.176, issue.1-3, 2003.
DOI : 10.1016/S0378-1127(02)00273-6

T. M. De-la-rosa, P. J. Aphalo, and T. Lehto, Effects of far-red light on the growth, mycorrhizas and mineral nutrition of Scots pine seedlings, Plant and Soil, vol.201, issue.1, pp.17-25, 1998.
DOI : 10.1023/A:1004383526878

T. M. De-la-rosa, T. Lehto, and P. J. Aphalo, Does far-red light affect growth and mycorrhizas of Scots pine seedlings grown in forest soil?, Plant and Soil, vol.211, issue.2, pp.259-268, 1999.
DOI : 10.1023/A:1004630524168

H. Dehlin, M. C. Nilsson, D. A. Wardle, and A. Shevtsova, Effects of shading and humus fertility on growth, competition, and ectomycorrhizal colonization of boreal forest tree seedlings, Canadian Journal of Forest Research, vol.34, issue.12, 2004.
DOI : 10.1139/x04-143

M. Dekker, M. Van-breugel, and F. J. Sterck, Effective height development of four co-occurring species in the gap-phase regeneration of Douglas fir monocultures under nature-oriented conversion, Forest Ecology and Management, vol.238, issue.1-3, pp.189-198, 2007.
DOI : 10.1016/j.foreco.2006.10.012

S. Delagrange, C. Messier, M. J. Lechowicz, and P. Dizengremel, Physiological, morphological and allocational plasticity in understory deciduous trees: importance of plant size and light availability, Tree Physiology, vol.24, issue.7, pp.775-784, 2004.
DOI : 10.1093/treephys/24.7.775

U. Dieguez-aranda, F. C. Dorado, J. G. Gonzadlez, and A. R. Alboreca, Dynamic growth model for Scots pine (Pinus sylvestris L.) plantations in Galicia (north-western Spain), Ecological Modelling, vol.191, issue.2, pp.225-242, 2006.
DOI : 10.1016/j.ecolmodel.2005.04.026

E. Dreyer, C. Collet, P. Montpied, and H. Sinoquet, Caract??risation de la tol??rance ?? l'ombrage de jeunes semis de H??tre et comparaison avec les essences associ??es., Revue Foresti??re Fran??aise, vol.57, issue.2, pp.175-188, 2005.
DOI : 10.4267/2042/5034

I. Ensminger, L. Schmidt, and J. Lloyd, Soil temperature and intermittent frost modulate the rate of recovery of photosynthesis in Scots pine under simulated spring conditions, New Phytologist, vol.172, issue.2, pp.428-442, 2008.
DOI : 10.1111/j.1469-8137.2007.02273.x

E. Fernbach and H. Mohr, Coaction of the blue/ultraviolet-A light and light absorbed by phytochrome in controlling growth of pine (Pinus sylvestris L.) seedlings, Planta, vol.180, pp.212-216, 1990.

C. Gamborg and J. B. Larsen, ???Back to nature??????a sustainable future for forestry?, Forest Ecology and Management, vol.179, issue.1-3, pp.559-571, 2003.
DOI : 10.1016/S0378-1127(02)00553-4

S. Gartner and A. Reif, The impact of forest transformation on stand structure and ground vegetation in the southern Black Forest, Germany, Plant and Soil, vol.264, issue.1/2, pp.35-51, 2004.
DOI : 10.1023/B:PLSO.0000047751.25915.77

J. Hansen, G. Vogg, and E. Beck, Assimilation, allocation and utilization of carbon by 3-year-old Scots pine (Pinus sylvestris L???.) trees during winter and early spring, Trees, vol.11, issue.2, pp.83-90, 1996.
DOI : 10.1007/PL00009659

M. Hyppönen, V. Alenius, and S. Valkonen, in Finnish Lapland, Scandinavian Journal of Forest Research, vol.75, issue.4, pp.347-357, 2005.
DOI : 10.1080/02827580410030154

C. M. Kaelke, E. L. Kruger, and P. B. Reich, Trade-offs in seedling survival, growth, and physiology among hardwood species of contrasting successional status along a light-availability gradient, Canadian Journal of Forest Research, vol.31, issue.9, pp.1602-1616, 2001.
DOI : 10.1139/x01-090

D. D. Kneeshaw, R. K. Kobe, K. D. Coates, and C. Messier, Sapling size influences shade tolerance ranking among southern boreal tree species, Journal of Ecology, vol.70, issue.2, pp.471-480, 2006.
DOI : 10.1046/j.1365-2745.2002.00691.x

R. K. Kobe, S. W. Pacala, J. A. Silander, and C. D. Canham, Juvenile Tree Survivorship as a Component of Shade Tolerance, Ecological Applications, vol.5, issue.2, pp.517-532, 1995.
DOI : 10.2307/1942040

T. T. Kozlowski, Physiological ecology of natural regeneration of harvested and disturbed forest stands: implications for forest management, Forest Ecology and Management, vol.158, issue.1-3, pp.195-221, 2002.
DOI : 10.1016/S0378-1127(00)00712-X

T. Kuuluvainen, Gap disturbance, ground microtopography, and the regeneration dynamics of boreal coniferous forests in Finland: a review, Ann. Zool. Fennici, vol.31, pp.35-51, 1994.

J. P. Schütz, Opportunities and strategies of transforming regular forests to irregular forests, Forest Ecology and Management, vol.151, issue.1-3, pp.87-94, 2001.
DOI : 10.1016/S0378-1127(00)00699-X

K. Seiwa, M. Ando, A. Imaji, M. Tomita, and K. Kanou, Spatio-temporal variation of environmental signals inducing seed germination in temperate conifer plantations and natural hardwood forests in northern Japan, Forest Ecology and Management, vol.257, issue.1, pp.361-369, 2009.
DOI : 10.1016/j.foreco.2008.09.002

G. Sonohat, P. Balandier, and F. Ruchaud, Predicting solar radiation transmittance in the understory of even-aged coniferous stands in temperate forests, Annals of Forest Science, vol.61, issue.7, pp.629-641, 2004.
DOI : 10.1051/forest:2004061
URL : https://hal.archives-ouvertes.fr/hal-00883798

S. Tanja, F. Berninger, T. Vesala, T. Markkanen, P. Hari et al., Air temperature triggers the recovery of evergreen boreal forest photosynthesis in spring, Global Change Biology, vol.21, issue.10, pp.1410-1426, 2003.
DOI : 10.1046/j.1365-2486.2000.00302.x

F. Valladares and U. Niinemets, Shade Tolerance, a Key Plant Feature of Complex Nature and Consequences, Annual Review of Ecology, Evolution, and Systematics, vol.39, issue.1, pp.237-257, 2008.
DOI : 10.1146/annurev.ecolsys.39.110707.173506

A. D. Vickers and S. C. Palmer, The influence of canopy cover and other factors upon the regeneration of Scots pine and its associated ground flora within Glen Tanar National Nature Reserve, Forestry, vol.73, issue.1, pp.37-49, 2000.
DOI : 10.1093/forestry/73.1.37

M. B. Walters and P. B. Reich, Low-light carbon balance and shade tolerance in the seedlings of woody plants: do winter deciduous and broad-leaved evergreen species differ?, New Phytologist, vol.143, issue.1, pp.143-154, 1999.
DOI : 10.1046/j.1469-8137.1999.00425.x

H. Williams, C. Messier, and D. D. Kneeshaw, Effects of light availability and sapling size on the growth and crown morphology of understory Douglas-fir and lodgepole pine, Canadian Journal of Forest Research, vol.29, issue.2, pp.222-231, 1999.
DOI : 10.1139/x98-189

N. Gaudio, P. Balandier, N. Donès, F. De-coligny, and C. Ginisty, RReShar (Regeneration and Resource Sharing), a model to simulate the uneven-aged forest stands dynamics, PARTIE 5 : paramétrage du modèle RReShar sous CAPSIS Présentation orale au colloque IUFRO « Uneven-Aged Forest Management, pp.23-26, 2010.

P. Adams, C. Beadle, N. Mendham, and P. Smethurst, The impact of timing and duration of grass control on growth of a young Eucalyptus globulus Labill, New Forests, vol.26, issue.2, pp.147-165, 2003.
DOI : 10.1023/A:1024490707175

R. Aerts, Aboveground Biomass and Nutrient Dynamics of Calluna vulgaris and Molinia caerulea in a Dry Heathland, Oikos, vol.56, issue.1, pp.31-38, 1989.
DOI : 10.2307/3566084

R. Aerts, R. Boot, and P. Vanderaart, The relation between above- and belowground biomass allocation patterns and competitive ability, Oecologia, vol.25, issue.4, pp.551-559, 1991.
DOI : 10.1007/BF00320419

M. Aikens, D. Ellum, J. Mckenna, M. Kelty, and M. Ashton, The effects of disturbance intensity on temporal and spatial patterns of herb colonization in a southern New England mixed-oak forest, Forest Ecology and Management, vol.252, issue.1-3, pp.144-158, 2007.
DOI : 10.1016/j.foreco.2007.06.031

T. Albaugh, H. Allen, B. Zutter, and H. Quicke, stands in the southeastern United States, Annals of Forest Science, vol.60, issue.7, pp.619-624, 2003.
DOI : 10.1051/forest:2003054
URL : https://hal.archives-ouvertes.fr/hal-00883734

R. Allain, S. Marie, F. Servol, J. Piroche, and T. Pérot, Les peuplements mélangés chêne-pin sylvestre de la forêt domaniale d'Orléans : caractérisation typologique. Essai d'utilisation des modèles de croissance en hauteur établis en futaie régulière pour l'évaluation de la fertilité des peuplements mélangés, 2004.

C. Ammer and S. Wagner, Problems and options in modelling fine-root biomass of single mature Norway spruce trees at given points from stand data, Canadian Journal of Forest Research, vol.32, issue.4, pp.581-590, 2002.
DOI : 10.1139/x01-229

P. Aphalo, C. Ballaré, and A. Scopel, Plant-plant signalling, the shade-avoidance response and competition, Journal of Experimental Botany, vol.50, issue.340, pp.1629-1634, 1999.
DOI : 10.1093/jxb/50.340.1629

I. Aranda, F. Pardo, L. Gil, and J. Pardos, Anatomical basis of the change in leaf mass per area and nitrogen investment with relative irradiance within the canopy of eight temperate tree species, Acta Oecologica, vol.25, issue.3, pp.187-195, 2004.
DOI : 10.1016/j.actao.2004.01.003

P. Aspe, Dynamique de la régénération naturelle chez le pin sylvestre, Pinus sylvestris L. : impacts de la structure des peuplements et des flux de gènes à différentes échelles spatiales sur la diversité génétique, Thèse (biologie forestière), 2009.

I. Aubin, M. Beaudet, and C. Messier, Light extinction coefficients specific to the understory vegetation of the southern boreal forest, Quebec, Canadian Journal of Forest Research, vol.30, issue.1, pp.168-177, 2000.
DOI : 10.1139/x99-185

D. Baize, Guide des analyses courantes en pédologie, INRA, vol.172, 1988.

P. Balandier and D. Pauwels, La lumière : outil sylvicole pour favoriser la diversité végétale ou la gestion cynégétique des peuplements de mélèze (Larix sp.), Forêt Wallonne, vol.61, pp.9-13, 2002.

P. Balandier, C. Collet, J. Miller, P. Reynolds, and S. Zedaker, Designing forest vegetation management strategies based on the mechanisms and dynamics of crop tree competition by neighbouring vegetation, Forestry, vol.79, issue.1, pp.3-27, 2006.
DOI : 10.1093/forestry/cpi056

P. Balandier, A. Marquier, N. Gaudio, L. Wehrlen, E. Casella et al., Methods for describing light capture by understorey weeds in temperate forests: consequences for tree regeneration. Proceedings de la conférence "Forest Vegetation Management -Towards Environmental Sustainability, pp.73-75, 2009.
URL : https://hal.archives-ouvertes.fr/hal-00468643

P. Balandier, H. Frochot, and A. Sourisseau, Improvement of direct tree seeding with cover crops in afforestation: Microclimate and resource availability induced by vegetation composition, Forest Ecology and Management, vol.257, issue.8, pp.1716-1724, 2009.
DOI : 10.1016/j.foreco.2009.01.032
URL : https://hal.archives-ouvertes.fr/hal-00964513

P. Balandier, A. Marquier, S. Perret, C. Collet, and B. Courbaud, Comment estimer la lumière dans le sousbois forestier à partir des caractéristiques dendrométriques des peuplements ? Rendez-vous Techniques ONF 27, pp.52-58, 2010.

C. Ballaré, Keeping up with the neighbours: phytochrome sensing and other signalling mechanisms, Trends in Plant Science, vol.4, issue.3, pp.97-102, 1999.
DOI : 10.1016/S1360-1385(99)01383-7

A. Ballester, A. Vieitez, and E. Vieitez, Allelopathic potential ofErica vagans, calluna vulgaris, andDaboecia cantabrica, Journal of Chemical Ecology, vol.39, issue.5, pp.851-857, 1982.
DOI : 10.1007/BF00994785

I. Barbeito, M. Fortin, and I. Canellas, Response of pine natural regeneration to small-scale spatial variation in a managed Mediterranean mountain forest, Applied Vegetation Science, vol.13, issue.4, pp.488-503, 2009.
DOI : 10.1111/j.1654-109X.2009.01043.x

S. Barbier, F. Gosselin, and P. Balandier, Influence of tree species on understory vegetation diversity and mechanisms involved???A critical review for temperate and boreal forests, Forest Ecology and Management, vol.254, issue.1, pp.1-15, 2008.
DOI : 10.1016/j.foreco.2007.09.038
URL : https://hal.archives-ouvertes.fr/hal-00964824

S. Barbier, P. Balandier, and F. Gosselin, Effet de quelques traits des arbres sur la r??partition des eaux de pluie en for??ts tem-p??r??es et bor??ales ???synth??se bibliographique, Annals of Forest Science, vol.64, issue.8, p.602, 2009.
DOI : 10.1051/forest/2009041

F. Bazzaz, Plant-plant interactions in successional environments (eds) Perspectives on plant competition, pp.239-263, 1990.

M. Becker and G. Lévy, Le point sur l'??cologie compar??e du Ch??ne sessile et du Ch??ne p??doncule, Revue Foresti??re Fran??aise, vol.2, issue.2, pp.148-153, 1990.
DOI : 10.4267/2042/26055

M. Begon, J. Harper, and C. Townsend, Ecology: individuals, populations and communities, Blackwell science, vol.1068, 1996.

J. Bengtsson, S. Nilsson, A. Franc, and P. Menozzi, Biodiversity, disturbances, ecosystem function and management of European forests, Forest Ecology and Management, vol.132, issue.1, pp.39-50, 2000.
DOI : 10.1016/S0378-1127(00)00378-9

P. Berbigier and J. Bonnefond, Measurement and modelling of radiation transmission within a stand of maritime pine (Pinus pinaster Ait), Annales des Sciences Foresti??res, vol.52, issue.1, pp.23-42, 1995.
DOI : 10.1051/forest:19950103
URL : https://hal.archives-ouvertes.fr/hal-00882977

N. Bréda, R. Huc, A. Granier, and E. Dreyer, Temperate forest trees and stands under severe drought: a review of ecophysiological responses, adaptation processes and long-term consequences, Annals of Forest Science, vol.63, issue.6, pp.625-644, 2006.
DOI : 10.1051/forest:2006042

A. Brêthes, Les types de station forestière de l'Orléanais, ONF, vol.400, 1993.

R. Brooker, D. Scott, S. Palmer, and E. Swaine, Transient facilitative effects of heather on Scots pine along a grazing disturbance gradient in Scottish moorland, Journal of Ecology, vol.78, issue.Suppl. 1, pp.637-645, 2006.
DOI : 10.1111/j.1365-2745.2006.01129.x

J. Buongiorno, J. Peyron, F. Houllier, and M. Bruciamacchie, Growth and management of mixed species, uneven-aged forests in the French Jura: implications for economic returns and tree diversity, Forest Science, vol.41, pp.397-429, 1995.
URL : https://hal.archives-ouvertes.fr/hal-01072893

R. Callaway, Positive interactions among plants, The Botanical Review, vol.43, issue.4, pp.306-349, 1995.
DOI : 10.1007/BF02912621
URL : https://hal.archives-ouvertes.fr/halsde-00295042

R. Callaway and L. Walker, COMPETITION AND FACILITATION: A SYNTHETIC APPROACH TO INTERACTIONS IN PLANT COMMUNITIES, Ecology, vol.78, issue.7, pp.1958-1965, 1997.
DOI : 10.2307/1939615

B. Casper and R. Jackson, PLANT COMPETITION UNDERGROUND, Annual Review of Ecology and Systematics, vol.28, issue.1, pp.545-570, 1997.
DOI : 10.1146/annurev.ecolsys.28.1.545

S. Catovsky and F. Bazzaz, Feedbacks between canopy composition and seedling regeneration in mixed conifer broad-leaved forests, Oikos, vol.127, issue.3, pp.403-420, 2002.
DOI : 10.1023/A:1009740016420

D. Chapman, M. Termansen, C. Quinn, N. Jin, A. Bonn et al., Modelling the coupled dynamics of moorland management and upland vegetation, Journal of Applied Ecology, vol.43, issue.2, pp.278-288, 2009.
DOI : 10.1111/j.1365-2664.2009.01618.x

Y. Claveau, C. Messier, P. Comeau, and K. Coates, Growth and crown morphological responses of boreal conifer seedlings and saplings with contrasting shade tolerance to a gradient of light and height, Canadian Journal of Forest Research, vol.32, issue.3, pp.458-468, 2002.
DOI : 10.1139/x01-220

Y. Claveau, C. Messier, and P. Comeau, Interacting influence of light and size on aboveground biomass distribution in sub-boreal conifer saplings with contrasting shade tolerance, Tree Physiology, vol.25, issue.3, pp.373-384, 2005.
DOI : 10.1093/treephys/25.3.373

B. Clinton, Light, temperature, and soil moisture responses to elevation, evergreen understory, and small canopy gaps in the southern Appalachians, Forest Ecology and Management, vol.186, issue.1-3, pp.243-255, 2003.
DOI : 10.1016/S0378-1127(03)00277-9

L. Coll, Compétition pour les ressources entre de jeunes hêtres (Fagus sylvatica L.) et la végétation dans des milieux en transition, Thèse (écologie forestière et écophysiologie), 2003.

L. Coll, P. Balandier, C. Picon-cochard, B. Prévosto, and C. T. , Competition for water between beech seedlings and surrounding vegetation in different light and vegetation composition conditions, Annals of Forest Science, vol.60, issue.7, pp.593-600, 2003.
DOI : 10.1051/forest:2003051
URL : https://hal.archives-ouvertes.fr/hal-00883731

L. Coll, P. Balandier, and C. Picon-cochard, Morphological and physiological responses of beech (Fagus sylvatica) seedlings to grass-induced belowground competition, Tree Physiology, vol.24, issue.1, pp.45-54, 2004.
DOI : 10.1093/treephys/24.1.45

J. Connell and R. Slatyer, Mechanisms of Succession in Natural Communities and Their Role in Community Stability and Organization, The American Naturalist, vol.111, issue.982, pp.1119-1144, 1977.
DOI : 10.1086/283241

J. Connell, Apparent versus "real" competition in plants (eds) Perspectives on plant competition, pp.9-26, 1990.

D. Coomes and P. Grubb, IMPACTS OF ROOT COMPETITION IN FORESTS AND WOODLANDS: A THEORETICAL FRAMEWORK AND REVIEW OF EXPERIMENTS, Ecological Monographs, vol.70, issue.2, pp.171-207, 2000.
DOI : 10.2307/1942248

P. Coquillard, Dynamique des systèmes agro-pastoraux de l'étage montagnard du Massif du Sancy et de la Chaîne des Puys : variations biologiques et fonctionnelles ; exemple d'application du modèle à Calluna vulgaris, Thèse, 1993.

B. Courbaud, Modélisation de la croissance en forêt irrégulière. Perspectives pour les pessières irrégulières de montagne. Revue Forestière Française n° spécial, pp.173-182, 1995.

C. Gamborg and J. Larsen, Back to nature' -a sustainable future for forestry? Forest Ecology and Management, pp.559-571, 2003.

E. Gardiner and J. Hodges, Growth and biomass distribution of cherrybark oak (Quercus pagoda Raf.) seedlings as influenced by light availability, Forest Ecology and Management, vol.108, issue.1-2, pp.127-134, 1998.
DOI : 10.1016/S0378-1127(98)00220-5

S. Gärtner and A. Reif, The impact of forest transformation on stand structure and ground vegetation in the southern Black Forest, Germany, Plant and Soil, vol.264, issue.1/2, pp.35-51, 2004.
DOI : 10.1023/B:PLSO.0000047751.25915.77

N. Gaudio, P. Balandier, and A. Marquier, Croissance de deux esp??ces comp??titrices (Rubus idaeus, Cytisus scoparius) colonisatrices des trou??es en for??t temp??r??e selon la disponibilit?? en lumi??re, Annals of Forest Science, vol.125, issue.1, p.104, 2008.
DOI : 10.1051/forest:2007076

N. Gaudio, Impact of adult trees on soil water content and consequences on regeneration Rapport de mission (COST-STSM-E47-4061), 2009.

N. Gaudio, P. Balandier, P. G. Dumas, Y. Jean, F. Ginisty et al., Light-mediated influence of three understorey species (Calluna vulgaris, Pteridium aquilinum, Molinia caerulea) on the growth of Pinus sylvestris seedlings, European Journal of Forest Research, vol.29, issue.1, pp.10342-10352, 1007.
DOI : 10.1007/s10342-010-0403-2
URL : https://hal.archives-ouvertes.fr/hal-00964576

L. George and F. Bazzaz, THE FERN UNDERSTORY AS AN ECOLOGICAL FILTER: EMERGENCE AND ESTABLISHMENT OF CANOPY-TREE SEEDLINGS, Ecology, vol.80, issue.3, pp.833-845, 1999.
DOI : 10.2307/1940078

J. Ghorbani, L. Duc, M. Mcallister, H. Pakeman, R. Marrs et al., Effects of the litter layer of Pteridium aquilinum on seed banks under experimental restoration, Applied Vegetation Science, vol.21, issue.65, pp.127-136, 2006.
DOI : 10.1111/j.1654-109X.2006.tb00662.x

M. Gillard, Rubus fruticosus) en fonction de la disponibilité en lumière dans le sous-bois forestier, 2010.

C. Gimingham, Calluna Salisb., The Journal of Ecology, vol.48, issue.2, pp.455-483, 1960.
DOI : 10.2307/2257528

S. Gliessman and C. Muller, The allelopathic mechanisms of dominance in bracken (Pteridium aquilinum) in Southern California, Journal of Chemical Ecology, vol.70, issue.3, pp.337-362, 1978.
DOI : 10.1007/BF00989342

D. Goldberg, Components of resource competition in plant communities (eds) Perspectives on plant competition, pp.27-49, 1990.

J. Grace, On the relationship between plant traits and competitive ability (eds) Perspectives on plant competition, pp.51-65, 1990.

A. Granier, N. Breda, P. Biron, and S. Villette, A lumped water balance model to evaluate duration and intensity of drought constraints in forest stands, Ecological Modelling, vol.116, issue.2-3, pp.269-283, 1999.
DOI : 10.1016/S0304-3800(98)00205-1

J. Grime, J. Hodgson, and R. Hunt, Comparative plant ecology, a functional approach to common British species, 2007.

M. Hancock, S. Egan, R. Summers, N. Cowie, A. Amphlett et al., The effect of experimental prescribed fire on the establishment of Scots pine Pinus sylvestris seedlings on heather Calluna vulgaris moorland, Forest Ecology and Management, vol.212, issue.1-3, pp.199-213, 2005.
DOI : 10.1016/j.foreco.2005.03.039

R. Hangs, J. Knight, V. Rees, and K. , Interspecific competition for nitrogen between early successional species and planted white spruce and jack pine seedlings, Canadian Journal of Forest Research, vol.32, issue.10, pp.1813-1821, 2002.
DOI : 10.1139/x02-100

R. Harmer and M. Robertson, Seedling root growth of six broadleaved tree species grown in competition with grass under irrigated nursery conditions, Annals of Forest Science, vol.60, issue.7, pp.601-608, 2003.
DOI : 10.1051/forest:2003052
URL : https://hal.archives-ouvertes.fr/hal-00883732

R. Harmer and G. Morgan, Development of Quercus robur advance regeneration following canopy reduction in an oak woodland, Forestry, vol.80, issue.2, pp.137-149, 2007.
DOI : 10.1093/forestry/cpm006

J. Harper, Population biology of plants, 1977.

H. Hasenauer, Concepts within tree growth modeling Sustainable forest management: growth models for Europe, pp.3-17, 2006.

H. Hasenauer and G. Kindermann, Methods for assessing regeneration establishment and height growth in uneven-aged mixed species stands, Forestry, vol.75, issue.4, pp.385-394, 2002.
DOI : 10.1093/forestry/75.4.385

B. Hawkins and G. Henry, Effect of Nitrogen Supply and Irradiance on Seedling Survival and Biomass in Two Evergreen, Ericaceous Species, Scandinavian Journal of Forest Research, vol.18, issue.5, pp.415-423, 2004.
DOI : 10.1016/S0378-1127(97)00285-5

S. Heinrichs and W. Schmidt, Short-term effects of selection and clear cutting on the shrub and herb layer vegetation during the conversion of even-aged Norway spruce stands into mixed stands, Forest Ecology and Management, vol.258, issue.5, pp.667-678, 2009.
DOI : 10.1016/j.foreco.2009.04.037

T. Heithecker and C. Halpern, Variation in microclimate associated with dispersed-retention harvests in coniferous forests of western Washington, Forest Ecology and Management, vol.226, issue.1-3, pp.60-71, 2006.
DOI : 10.1016/j.foreco.2006.01.024

J. Holeksa, Relationship between field-layer vegetation and canopy openings in a Carpathian subalpine spruce forest, Plant Ecology, vol.168, issue.1, pp.57-67, 2003.
DOI : 10.1023/A:1024457303815

J. Humphrey and M. Swaine, Factors affecting the natural regeneration of Quercus in Scottish oakwoods .1. Competition from Pteridium aquilinum, Journal of Applied Ecology, vol.34, pp.577-584, 1997.

M. Hyppönen, V. Alenius, and S. Valkonen, in Finnish Lapland, Scandinavian Journal of Forest Research, vol.75, issue.4, pp.347-357, 2005.
DOI : 10.1080/02827580410030154

J. Hytönen and P. Jylhä, Effects of competing vegetation and post-planting weed control on the mortality, growth and vole damages to Betula pendula planted on former agricultural land, Silva Fennica, vol.39, issue.3, pp.365-380, 2005.
DOI : 10.14214/sf.374

S. Iivonen, R. Rikala, and E. Vapaavuori, Seasonal root growth of Scots pine seedlings in relation to shoot phenology, carbohydrate status, and nutrient supply, Canadian Journal of Forest Research, vol.31, issue.9, pp.1569-1578, 2001.
DOI : 10.1139/x01-094

M. Jach and R. Ceulemans, Effects of elevated atmospheric CO2 on phenology, growth and crown structure of Scots pine (Pinus sylvestris) seedlings after two years of exposure in the field, Tree Physiology, vol.19, issue.4-5, pp.289-300, 1999.
DOI : 10.1093/treephys/19.4-5.289

M. Jalal, D. Read, and E. Haslam, Phenolic composition and its seasonal variation in Calluna vulgaris, Phytochemistry, vol.21, issue.6, pp.1397-1401, 1982.
DOI : 10.1016/0031-9422(82)80150-7

S. Janecek and J. Leps, Effect of litter, leaf cover and cover of basal internodes of the dominant species Molinia caerulea on seedling recruitment and established vegetation, Acta Oecologica, vol.28, issue.2, pp.141-147, 2005.
DOI : 10.1016/j.actao.2005.03.006

T. Jefferies, Ecology of the Purple Heath Grass (Molinia Caerulea), The Journal of Ecology, vol.3, issue.2, pp.93-109, 1915.
DOI : 10.2307/2962376

R. Jobidon, Density-dependent effects of northern hardwood competition on selected environmental resources and young white spruce (Picea glauca) plantation growth, mineral nutrition, and stand structural development ??? a 5-year study, Forest Ecology and Management, vol.130, issue.1-3, pp.77-97, 2000.
DOI : 10.1016/S0378-1127(99)00176-0

C. Kaelke, E. Kruger, and P. Reich, Trade-offs in seedling survival, growth, and physiology among hardwood species of contrasting successional status along a light-availability gradient, Canadian Journal of Forest Research, vol.31, issue.9, pp.1602-1616, 2001.
DOI : 10.1139/x01-090

C. Karlsson and G. Orlander, Soil Scarification Shortly before a Rich Seed Fall Improves Seedling Establishment in Seed Tree Stands of Pinus sylvestris, Scandinavian Journal of Forest Research, vol.15, issue.2, pp.256-266, 2000.
DOI : 10.1080/028275800750015073

P. Keddy, Competitive hierarchies and centrifugal organization in plant communities (eds) Perspectives on plant competition, pp.265-290, 1990.

D. King, Allocation of above-ground growth is related to light in temperate deciduous saplings, Functional Ecology, vol.7, issue.4, pp.482-488, 2003.
DOI : 10.1046/j.1469-8137.1999.00425.x

D. Kneeshaw, R. Kobe, K. Coates, and C. Messier, Sapling size influences shade tolerance ranking among southern boreal tree species, Journal of Ecology, vol.70, issue.2, pp.471-480, 2006.
DOI : 10.1046/j.1365-2745.2002.00691.x

R. Kobe, S. Pacala, J. Silander, and C. Canham, Juvenile Tree Survivorship as a Component of Shade Tolerance, Ecological Applications, vol.5, issue.2, pp.517-532, 1995.
DOI : 10.2307/1942040

H. Koop and P. Hilgen, Forest dynamics and regeneration mosaic shifts in unexploited beech (Fagus sylvatica) stands at Fontainebleau (France), Forest Ecology and Management, vol.20, issue.1-2, pp.135-150, 1987.
DOI : 10.1016/0378-1127(87)90155-1

T. Kozlowski, Control of shoot growth Physiological ecology, growth and development of trees, pp.296-386, 1971.

T. Kozlowski, Physiological ecology of natural regeneration of harvested and disturbed forest stands: implications for forest management, Forest Ecology and Management, vol.158, issue.1-3, pp.195-221, 2002.
DOI : 10.1016/S0378-1127(00)00712-X

G. Kunstler, Dynamique du paysage et démographie des espèces arborées dominantes. La régénération du hêtre et du chêne sur les Grands Causses, Thèse (biologie des populations et écologie), ENSAM, 2005.

G. Kunstler, T. Curt, M. Bouchaud, and J. Lepart, Indirect facilitation and competition in tree species colonization of sub-Mediterranean grasslands, Journal of Vegetation Science, vol.54, issue.3, pp.379-388, 2006.
DOI : 10.1111/j.1654-1103.2006.tb02458.x

T. Kuuluvainen, Gap disturbance, ground microtopography, and the regeneration dynamics of boreal coniferous forests in Finland: a review, Ann. Zool. Fennici, vol.31, pp.35-51, 1994.

R. Lautenschlager, Environmental resource interactions affect red raspberry growth and its competition with white spruce, Canadian Journal of Forest Research, vol.29, issue.7, pp.906-916, 1999.
DOI : 10.1139/x99-078

L. Duc, M. Pakeman, R. Marrs, and R. , A restoration experiment on moorland infested by Pteridium aquilinum: Plant species responses, Agriculture, Ecosystems & Environment, vol.119, issue.1-2, pp.53-59, 2007.
DOI : 10.1016/j.agee.2006.06.008

J. Levine, INDIRECT FACILITATION: EVIDENCE AND PREDICTIONS FROM A RIPARIAN COMMUNITY, Ecology, vol.80, issue.5, pp.1762-1769, 1999.
DOI : 10.2307/1940449

V. Lieffers, C. Messier, K. Stadt, F. Gendron, and P. Comeau, Predicting and managing light in the understory of boreal forests, Canadian Journal of Forest Research, vol.29, issue.6, pp.796-811, 1999.
DOI : 10.1139/x98-165

M. Löf, Establishment and growth in seedlings of <i>Fagus sylvatica</i> and <i>Quercus robur</i>: influence of interference from herbaceous vegetation, Canadian Journal of Forest Research, vol.30, issue.6, pp.855-864, 2000.
DOI : 10.1139/cjfr-30-6-855

M. Löf and N. Welander, L., Annals of Forest Science, vol.61, issue.8, pp.781-788, 2004.
DOI : 10.1051/forest:2004075
URL : https://hal.archives-ouvertes.fr/in2p3-00276253

M. Löf, A. Bolte, and N. Welander, seedlings, Scandinavian Journal of Forest Research, vol.17, issue.4, pp.322-328, 2005.
DOI : 10.1016/S0378-1127(97)00326-5
URL : https://hal.archives-ouvertes.fr/hal-00883438

M. Löf, J. Bergquist, J. Brunet, M. Karlsson, and T. Welander, Conversion of Norway spruce to broadleaves regeneration systems, fencing and performance of planted seedlings, Ecological Bulletins, vol.53, 2010.

H. Lyr and V. Garbe, Influence of root temperature on growth of Pinus sylvestris, Fagus sylvatica, Tilia cordata and Quercus robur, Trees, vol.9, issue.4, pp.220-223, 1995.
DOI : 10.1007/BF00195276

H. Lyr, Effect of the root temperature on growth parameters of various European tree species, Annales des Sciences Foresti??res, vol.53, issue.2-3, pp.317-323, 1996.
DOI : 10.1051/forest:19960214
URL : https://hal.archives-ouvertes.fr/hal-00883054

D. Malcolm, W. Mason, and G. Clarke, The transformation of conifer forests in Britain ??? regeneration, gap size and silvicultural systems, Forest Ecology and Management, vol.151, issue.1-3, pp.7-23, 2001.
DOI : 10.1016/S0378-1127(00)00692-7

A. Mallik, Conifer Regeneration Problems in Boreal and Temperate Forests with Ericaceous Understory: Role of Disturbance, Seedbed Limitation, and Keytsone Species Change, Critical Reviews in Plant Sciences, vol.22, issue.3-4, pp.341-366, 2003.
DOI : 10.1080/713610860

R. Marrs and A. Watt, Biological Flora of the British Isles: Pteridium aquilinum (L.) Kuhn, Journal of Ecology, vol.41, issue.37, pp.1272-1321, 2006.
DOI : 10.1006/anbo.1999.1068

W. Mason, C. Edwards, and S. Hale, Survival and early seedling growth of conifers with different shade tolerance in a Sitka spruce spacing trial and relationship to understorey light climate, Silva Fennica, vol.38, issue.4, pp.357-370, 2004.
DOI : 10.14214/sf.404

P. Mayer, Hot Spot: Forest policy in Europe: achievements of the MCPFE and challenges ahead, Forest Policy and Economics, vol.1, issue.2, pp.177-185, 2000.
DOI : 10.1016/S1389-9341(00)00018-6

N. Mccarthy, N. Bentsen, I. Willoughby, and P. Balandier, The state of forest vegetation management in Europe in the 21st century, European Journal of Forest Research, vol.72, issue.2, pp.10342-10352, 1007.
DOI : 10.1007/s10342-010-0429-5
URL : https://hal.archives-ouvertes.fr/hal-00964764

F. Medail, Biogéographie, écologie et valeur patrimoniale des forêts de pin sylvestre (Pinus sylvestris L.) en région méditerranéenne, Forêt Méditerranéenne, vol.22, pp.5-22, 2001.

C. Messier, Effects of neutral shade and growing media on growth, biomass allocation, and competitive ability of Gaultheria shallon, Canadian Journal of Botany, vol.70, pp.271-2276, 1992.

C. Messier and P. Puttonen, Spatial and temporal variation in the Bight environment of developing Scots pine stands: the basis for a quick and efficient method of characterizing Bight, Canadian Journal of Forest Research, vol.25, issue.2, pp.343-354, 1995.
DOI : 10.1139/x95-038

C. Messier, R. Doucet, J. Ruel, Y. Claveau, C. Kelly et al., Functional ecology of advance regeneration in relation to light in boreal forests, Canadian Journal of Forest Research, vol.29, issue.6, pp.812-823, 1999.
DOI : 10.1139/x99-070

J. Miina, K. Eerikäinen, and H. Hasenauer, Modeling forest regeneration Sustainable forest management: growth models for Europe, pp.93-109, 2006.

R. Mitchell, R. Rose, and S. Palmer, Restoration of Calluna vulgaris on grass-dominated moorlands: The importance of disturbance, grazing and seeding, Biological Conservation, vol.141, issue.8, pp.2100-2111, 2008.
DOI : 10.1016/j.biocon.2008.06.006

F. Moola and A. Mallik, Morphological plasticity and regeneration strategies of velvet leaf blueberry (Vaccinium myrtilloides Michx.) following canopy disturbance in boreal mixedwood forests, Forest Ecology and Management, vol.111, issue.1, pp.35-50, 1998.
DOI : 10.1016/S0378-1127(98)00306-5

F. Morneau, C. Duprez, and J. Hervé, Les forêts mélangées en France métropolitaine. Caractérisation à partir des résultats de l, Inventaire Forestier National. Revue forestière française, vol.2, pp.107-120, 2008.

G. Nabuurs, Quantification of herb layer dynamics under tree canopy, Forest Ecology and Management, vol.88, issue.1-2, pp.143-148, 1996.
DOI : 10.1016/S0378-1127(96)03820-0

E. Nambiar, Interplay between nutrients, water, root growth and productivity in young plantations, Forest Ecology and Management, vol.30, issue.1-4, pp.213-232, 1990.
DOI : 10.1016/0378-1127(90)90138-2

E. Nambiar and R. Sands, Competition for water and nutrients in forests, Canadian Journal of Forest Research, vol.23, issue.10, pp.1955-1968, 1993.
DOI : 10.1139/x93-247

N. Bieng and M. , Construction de modèles de structure spatiale permettant de simuler des peuplements virtuels réalistes. Application aux peuplements mélangés chêne sessile -pin sylvestre de la région Centre, Thèse (sciences forestières), ENGREF, 2007.

U. Niinemets, GLOBAL-SCALE CLIMATIC CONTROLS OF LEAF DRY MASS PER AREA, DENSITY, AND THICKNESS IN TREES AND SHRUBS, Ecology, vol.82, issue.2, pp.453-469, 2001.
DOI : 10.1139/x93-199

U. Niinemets, The controversy over traits conferring shade-tolerance in trees: ontogenetic changes revisited, Journal of Ecology, vol.81, issue.2, pp.464-470, 2006.
DOI : 10.1038/nature02403

M. Nilsson and D. Wardle, Understory vegetation as a forest ecosystem driver: evidence from the northern Swedish boreal forest, Frontiers in Ecology and the Environment, vol.3, issue.8, pp.421-428, 2005.
DOI : 10.1890/1540-9295(2005)003[0421:UVAAFE]2.0.CO;2

G. Norberg, A. Dolling, A. Jaderlund, M. Nilsson, and O. Zackrisson, Control of heather (Calluna vulgaris (L.) Hull) by steam treatment: Effects on establishment and early growth of Scots pine, pp.187-198, 2001.

O. Nystrand and A. Granström, Forest floor moisture controls predator activity on juvenile seedlings of Pinus sylvestris, Canadian Journal of Forest Research, vol.27, issue.11, pp.1746-1752, 1997.
DOI : 10.1139/x97-148

T. Oleksyn, P. Reich, M. Tjoelker, and W. Chalupka, Biogeographic differences in shoot elongation pattern among European Scots pine populations, Forest Ecology and Management, vol.148, issue.1-3, pp.207-220, 2001.
DOI : 10.1016/S0378-1127(00)00537-5

B. Oyen, H. Blom, I. Gjerde, T. Myking, M. Saetersdal et al., Ecology, history and silviculture of Scots pine (Pinus sylvestris L.) in western Norway - a literature review, Forestry, vol.79, issue.3, pp.319-329, 2006.
DOI : 10.1093/forestry/cpl019

S. Pacala, C. Canham, J. Silander, and R. Kobe, Sapling growth as a function of resources in a north temperate forest, Canadian Journal of Forest Research, vol.24, issue.11, pp.2172-2183, 1994.
DOI : 10.1139/x94-280

S. Pacala, C. Canham, J. Saponara, J. Silander, R. Kobe et al., Forest Models Defined by Field Measurements: Estimation, Error Analysis and Dynamics, Ecological Monographs, vol.66, issue.1, pp.1-43, 1996.
DOI : 10.2307/2963479

J. Pagès and R. Michalet, A test of the indirect facilitation model in a temperate hardwood forest of the northern French Alps, Journal of Ecology, vol.8, issue.6, pp.932-940, 2003.
DOI : 10.1016/0169-5347(90)90095-U

J. Pagès and R. Michalet, Contrasted Responses of Two Understorey Species to Direct and Indirect Effects of a Canopy Gap, Plant Ecology, vol.67, issue.2, pp.179-187, 2006.
DOI : 10.1007/s11258-005-0976-x

J. Paluch, The influence of the spatial pattern of trees on forest floor vegetation and silver fir (Abies alba Mill.) regeneration in uneven-aged forests, Forest Ecology and Management, vol.205, issue.1-3, pp.283-298, 2005.
DOI : 10.1016/j.foreco.2004.10.010

M. Pardos, F. Montes, I. Aranda, and I. Canellas, Influence of environmental conditions on germinant survival and diversity of Scots pine (Pinus sylvestris L.) in central Spain, European Journal of Forest Research, vol.140, issue.1, pp.37-47, 2007.
DOI : 10.1007/s10342-005-0090-6

W. Parker, D. Pitt, and A. Morneault, Influence of woody and herbaceous competition on microclimate and growth of eastern white pine (Pinus strobus L.) seedlings planted in a central Ontario clearcut, Forest Ecology and Management, vol.258, issue.9, pp.2013-2025, 2009.
DOI : 10.1016/j.foreco.2009.07.046

T. Pérot, S. Perret, C. Meredieu, and C. Ginisty, Pr??voir la croissance et la production du Pin sylvestre : le module Sylvestris sous Capsis 4, Revue Foresti??re Fran??aise, vol.1, issue.1, pp.57-84, 2007.
DOI : 10.4267/2042/8483

T. Pérot, F. Goreaud, and C. Ginisty, Quels mod??les de croissance pour les peuplements m??lang??s ? Exemple du m??lange Ch??ne sessile - Pin sylvestre, Revue Foresti??re Fran??aise, vol.2, issue.2, pp.215-232, 2008.
DOI : 10.4267/2042/18145

T. Pérot, Quel est le niveau de détail pertinent pour modéliser la croissance d'une forêt mélangée ? Comparaison d'une famille de modèles et application aux peuplements mélangés chêne sessile -pin sylvestre, 2009.

A. Petritan, V. Lüpke, B. Petritan, and I. , Influence of light availability on growth, leaf morphology and plant architecture of beech (Fagus sylvatica L.), maple (Acer pseudoplatanus L.) and ash (Fraxinus excelsior L.) saplings, European Journal of Forest Research, vol.77, issue.39, pp.61-74, 2009.
DOI : 10.1007/s10342-008-0239-1

C. Picon-cochard, L. Coll, and P. Balandier, The role of below-ground competition during early stages of secondary succession: the case of 3-year-old Scots pine (Pinus sylvestris L.) seedlings in an abandoned grassland, Oecologia, vol.10, issue.3, pp.373-383, 2006.
DOI : 10.1007/s00442-006-0379-2

A. Porté and H. Bartelink, Modelling mixed forest growth: a review of models for forest management, Ecological Modelling, vol.150, issue.1-2, pp.141-188, 2002.
DOI : 10.1016/S0304-3800(01)00476-8

A. Portsmuth and U. Niinemets, Structural and physiological plasticity in response to light and nutrients in five temperate deciduous woody species of contrasting shade tolerance, Functional Ecology, vol.25, issue.1, pp.61-77, 2007.
DOI : 10.1111/j.1365-2435.2006.01208.x

A. Pouzerat, Répartition horizontale et verticale des racines d'arbres et de la végétation du sous-bois en peuplement mélangé chêne sessile-pin sylvestre; influence sur la teneur en eau, Université de Bordeaux 1 (France), 2010.

B. Prévosto, Analyse fonctionnelle des boisements spontanés et modélisation de leur développement Application aux peuplements de Betula pendula (Roth.) et Pinus sylvestris (L.) en moyenne montagne volcanique (la chaîne des Puys, Thèse (écologie), 1999.

B. Prévosto, T. Curt, J. Gueugnot, and P. Coquillard, Modeling mid-elevation Scots pine growth on a volcanic substrate, Forest Ecology and Management, vol.131, issue.1-3, pp.223-237, 2000.
DOI : 10.1016/S0378-1127(99)00216-9

B. Prévosto, A. Robert, and P. Coquillard, Development of Cytisus scoparius L. at stand and individual level in a mid-elevation mountain of the French Massif Central, Acta Oecologica, vol.25, issue.1-2, pp.73-81, 2004.
DOI : 10.1016/j.actao.2003.11.005

B. Prévosto, E. Dambrine, P. Coquillard, and A. Robert, Broom (Cytisus??scoparius) colonization after grazing abandonment in??the??French Massif Central: impact on??vegetation composition and??resource availability, Acta Oecologica, vol.30, issue.2, pp.258-268, 2006.
DOI : 10.1016/j.actao.2006.05.001

D. Price, N. Zimmermann, P. Van-der-meer, M. Lexer, P. Leadley et al., Regeneration in gap models: priority issues for studying forest responses to climate change, Climatic Change, vol.51, issue.3/4, pp.475-508, 2001.
DOI : 10.1023/A:1012579107129

J. Pritchard and P. Comeau, Effects of opening size and stand characteristics on light transmittance and temperature under young trembling aspen stands, Forest Ecology and Management, vol.200, issue.1-3, pp.119-128, 2004.
DOI : 10.1016/j.foreco.2004.06.002

D. Provendier and P. Balandier, Comparaison des effets de la comp??tition par les gramin??es (Gramino??des) et le gen??t (Cytisus scoparius) sur la croissance et les traits fonctionnels de plants de h??tre (Fagus sylvatica), Annals of Forest Science, vol.29, issue.5, p.510, 2008.
DOI : 10.1051/forest:2008028

J. Rameau, D. Mansion, and G. Dume, Flore forestière française, Plaines et collines. IDF, vol.1, issue.1794, 1989.

J. Rameau, Comportement dynamique du Ch??ne p??doncul?? et du Ch??ne sessile dans les successions foresti??res, Revue Foresti??re Fran??aise, vol.2, issue.2, pp.155-164, 1990.
DOI : 10.4267/2042/26056

J. Rameau, Accrus, successions végétales et modèles de dynamique linéaire forestière. Ingénieries n°spécial "Boisements naturels des espaces agricoles en déprise, pp.33-48, 1999.

P. Reich, M. Tjoelker, M. Walters, D. Vanderklein, and C. Buschena, Close association of RGR, leaf and root morphology, seed mass and shade tolerance in seedlings of nine boreal tree species grown in high and low light, Functional Ecology, vol.12, issue.3, pp.327-338, 1998.
DOI : 10.1086/285088

E. Ribbens, J. Silander, and S. Pacala, Seedling Recruitment in Forests: Calibrating Models to Predict Patterns of Tree Seedling Dispersion, Ecology, vol.75, issue.6, pp.1794-1806, 1994.
DOI : 10.2307/1939638

J. Ricard and C. Messier, Abundance, growth and allometry of red raspberry (Rubus idaeus L.) along a natural light gradient in a northern hardwood forest, Forest Ecology and Management, vol.81, issue.1-3, pp.153-160, 1996.
DOI : 10.1016/0378-1127(95)03643-1

J. Ricard, C. Messier, S. Delagrange, and M. Beaudet, Do understory sapling respond to both light and below-ground competition?: a field experiment in a north-eastern American hardwood forest and a literature review, Annals of Forest Science, vol.60, issue.8, pp.749-756, 2003.
DOI : 10.1051/forest:2003069
URL : https://hal.archives-ouvertes.fr/hal-00883749

D. Richardson, Ecology and biogeography of Pinus, 2000.

R. Robinson, The Production by Roots of Calluna Vulgaris of a Factor Inhibitory to Growth of some Mycorrhizal Fungi, The Journal of Ecology, vol.60, issue.1, pp.219-224, 1972.
DOI : 10.2307/2258051

T. Robson, J. Rodriguez-calcerrada, D. Sanchez-gomez, and I. Aranda, Summer drought impedes beech seedling performance more in a sub-Mediterranean forest understory than in small gaps, Tree Physiology, vol.29, issue.2, pp.249-259, 2009.
DOI : 10.1093/treephys/tpn023

J. Rodriguez-calcerrada, S. Mutke, A. J. Gil, L. Pardos, J. Aranda et al., Influence of overstory density on understory light, soil moisture, and survival of two underplanted oak species in a Mediterranean montane Scots pine forest, Investigaci??n Agraria: Sistemas y Recursos Forestales, vol.17, issue.1, pp.31-38, 2008.
DOI : 10.5424/srf/2008171-01021

J. Rodriguez-calcerrada, F. Cano, M. Valbuena-carabana, L. Gil, and I. Aranda, Functional performance of oak seedlings naturally regenerated across microhabitats of distinct overstorey canopy closure, New Forests, vol.81, issue.2, pp.245-259, 2010.
DOI : 10.1007/s11056-009-9168-1

A. Royo and W. Carson, On the formation of dense understory layers in forests worldwide: consequences and implications for forest dynamics, biodiversity, and succession, Canadian Journal of Forest Research, vol.36, issue.6, pp.1345-1362, 2006.
DOI : 10.1139/x06-025

W. Rundel and B. Yoder, Ecophysiolohy of Pinus Ecology and biogeography of Pinus, pp.296-323, 2000.

K. Rünk and K. Zobel, Phenotypic plasticity and biomass allocation pattern in three Dryopteris (Dryopteridaceae) species on an experimental light-availability gradient, Plant Ecology, vol.9, issue.1, pp.85-99, 2007.
DOI : 10.1007/s11258-006-9250-0

L. Sack and P. Grubb, Why do species of woody seedlings change rank in relative growth rate between low and high irradiance?, Functional Ecology, vol.11, issue.2, pp.145-154, 2001.
DOI : 10.1139/cjfr-28-6-871

D. Sanchez-gomez, M. Zavala, and F. Valladares, Seedling survival responses to??irradiance are differentially influenced by??low-water availability in??four??tree species of??the??Iberian cool temperate???Mediterranean ecotone, Acta Oecologica, vol.30, issue.3, pp.322-332, 2006.
DOI : 10.1016/j.actao.2006.05.005

P. Schütt, W. Koch, H. Blaschke, K. Lang, H. Schuck et al., Avant que forêt ne meure : le dépérissement de la forêt -principaux dégâts observés et évolution, Delachaux et Niestlé, vol.96, 1984.

J. Schütz, Close-to-nature silviculture: is this concept compatible with species diversity?, Forestry, vol.72, issue.4, pp.359-366, 1999.
DOI : 10.1093/forestry/72.4.359

J. Schütz, Opportunities and strategies of transforming regular forests to irregular forests, Forest Ecology and Management, vol.151, issue.1-3, pp.87-94, 2001.
DOI : 10.1016/S0378-1127(00)00699-X

J. Schütz, Opportunistic methods of controlling vegetation, inspired by natural plant succession dynamics with special reference to natural outmixing tendencies in a gap regeneration, Annals of Forest Science, vol.61, issue.2, pp.149-156, 2004.
DOI : 10.1051/forest:2004006

C. Shropshire, R. Wagner, F. Bell, and C. Swanton, Light attenuation by early successional plants of the boreal forest, Canadian Journal of Forest Research, vol.31, issue.5, pp.812-823, 2001.
DOI : 10.1139/x01-006

J. Silbernagel and M. Moeur, Modeling canopy openness and understory gap patterns based on image analysis and mapped tree data, Forest Ecology and Management, vol.149, issue.1-3, pp.217-233, 2001.
DOI : 10.1016/S0378-1127(00)00556-9

G. Sonohat, H. Sinoquet, C. Varler-grancher, M. Rakocevic, A. Jacquet et al., Leaf dispersion and light partitioning in three-dimensionally digitized tall fescue-white clover mixtures, Plant, Cell and Environment, vol.47, issue.4, pp.529-538, 2002.
DOI : 10.1006/anbo.1998.0843

G. Sonohat, P. Balandier, H. Sinoquet, C. Varlet-grancher, R. Jouvie et al., Mesure de l'éclairement au sol dans les peuplements forestiers réguliers et relations avec leurs caractéristiques dendrométriques, 2003.

K. Stadt and V. Lieffers, Comparing PAR transmission models for forest understorey vegetation, Applied Vegetation Science, vol.47, issue.1, pp.65-76, 2005.
DOI : 10.1111/j.1654-109X.2005.tb00630.x

K. Taylor, A. Rowland, and H. Jones, Molinia caerulea (L.) Moench, Journal of Ecology, vol.68, issue.1, pp.126-144, 2001.
DOI : 10.1046/j.1365-2745.2000.00455.x

S. Thomas, C. Halpern, D. Falk, D. Liguori, and K. Austin, PLANT DIVERSITY IN MANAGED FORESTS: UNDERSTORY RESPONSES TO THINNING AND FERTILIZATION, Ecological Applications, vol.9, issue.3, pp.864-879, 1999.
DOI : 10.2307/1939319

R. Thomsen, J. Svenning, and H. Balslev, Overstorey Control of Understorey Species Composition in a Near-natural Temperate Broadleaved Forest in Denmark, Plant Ecology, vol.5, issue.1, pp.113-126, 2005.
DOI : 10.1007/s11258-005-3996-7

D. Tilman, Plant strategies and the dynamics and structure of plant communities, 1988.

D. Tilman, Mechanisms of plant competition for nutrients: the elements of a predictive theory of competition Perspectives on plant competition, pp.117-141, 1990.

J. Timbal, J. Gelpe, and J. Garbaye, ??tude pr??liminaire sur l'effet d??pressif de la molinie (Molinia caerulea) sur la croissance et l'??tat mycorhizien de semis de ch??ne rouge (Quercus rubra), Annales des Sciences Foresti??res, vol.47, issue.6, pp.643-649, 1990.
DOI : 10.1051/forest:19900609

J. Timbal and G. Aussenac, An overview of ecology and silviculture of indigenous oaks in France, Annales des Sciences Foresti??res, vol.53, issue.2-3, pp.649-661, 1996.
DOI : 10.1051/forest:19960243
URL : https://hal.archives-ouvertes.fr/hal-00883083

F. Valladares, J. Chico, I. Aranda, L. Balaguer, P. Dizengremel et al., The greater seedling high-light tolerance of Quercus robur over Fagus sylvatica is linked to a greater physiological plasticity, Trees, vol.16, pp.395-403, 2002.

A. Van-hees, Growth and morphology of pedunculate oak (Quercus robur L) and beech (Fagus sylvatica L) seedlings in relation to shading and drought, Annales des Sciences Foresti??res, vol.54, issue.1, pp.9-18, 1997.
DOI : 10.1051/forest:19970102
URL : https://hal.archives-ouvertes.fr/hal-00883128

A. Van-hees and A. Clerkx, Shading and root???shoot relations in saplings of silver birch, pedunculate oak and beech, Forest Ecology and Management, vol.176, issue.1-3, pp.439-448, 2003.
DOI : 10.1016/S0378-1127(02)00307-9

A. Vickers and S. Palmer, The influence of canopy cover and other factors upon the regeneration of Scots pine and its associated ground flora within Glen Tanar National Nature Reserve, Forestry, vol.73, issue.1, pp.37-49, 2000.
DOI : 10.1093/forestry/73.1.37

V. Lüpke and B. , Silvicultural methods of oak regeneration with special respect to shade tolerant mixed species, Forest Ecology and Management, vol.106, issue.1, pp.19-26, 1998.
DOI : 10.1016/S0378-1127(97)00235-1

V. Teuffel, K. Hein, S. Kotar, M. Preuhsler, E. Puumalainen et al., End user needs and requirements Sustainable forest management: growth models for Europe, pp.19-38, 2006.

R. Wagner, K. Little, B. Richardson, and K. Mcnabb, The role of vegetation management for enhancing productivity of the world's forests, Forestry, vol.79, issue.1, pp.57-79, 2006.
DOI : 10.1093/forestry/cpi057

S. Wagner, H. Fisher, and F. Huth, Canopy effects on vegetationForest Vegetation Management -Towards Environmental Sustainability, Proceedings de la conférence, pp.52-54, 2009.

S. Wagner, H. Fisher, and F. Huth, Canopy effects on vegetation caused by harvesting and regeneration treatments, European Journal of Forest Research, vol.255, issue.4, 2010.
DOI : 10.1007/s10342-010-0378-z

M. Walters and P. Reich, seedlings in deeply shaded understories of northern Wisconsin: effects of nitrogen and water availability, Canadian Journal of Forest Research, vol.27, issue.2, pp.237-247, 1997.
DOI : 10.1139/x96-178

M. Walters and P. Reich, Low-light carbon balance and shade tolerance in the seedlings of woody plants: do winter deciduous and broad-leaved evergreen species differ?, New Phytologist, vol.143, issue.1, pp.143-154, 1999.
DOI : 10.1046/j.1469-8137.1999.00425.x

P. Weber, H. Bugmann, and A. Rigling, Radial growth responses to drought of Pinus sylvestris and Quercus pubescens in an inner-Alpine dry valley, Journal of Vegetation Science, vol.57, issue.6, pp.777-792, 2007.
DOI : 10.1111/j.1654-1103.2007.tb02594.x

P. Weisberg, C. Hadorn, and H. Bugmann, Beziehungen zwischen Bodenvegetation und Bestandeseigenschaften in zwei Gebirgsw???ldern der gem???ssigten Zone, Forstwissenschaftliches Centralblatt, vol.122, issue.5, pp.273-286, 2003.
DOI : 10.1007/s10342-003-0004-4

H. Williams, C. Messier, and D. Kneeshaw, Effects of light availability and sapling size on the growth and crown morphology of understory Douglas-fir and lodgepole pine, Canadian Journal of Forest Research, vol.29, issue.2, pp.222-231, 1999.
DOI : 10.1139/x98-189

I. Willoughby, Future alternatives to the use of herbicides in British forestry, Canadian Journal of Forest Research, vol.29, issue.7, pp.866-874, 1999.
DOI : 10.1139/x99-043

I. Willoughby, P. Balandier, N. Bentsen, N. Mccarthy, and J. Claridge, Forest vegetation in Europe: current practice and future requirements. Rapport final de la Cost Action E47, Hampshire, vol.156, 2009.
URL : https://hal.archives-ouvertes.fr/hal-00468013

R. Zweifel, A. Rigling, and M. Dobbertin, Species-specific stomatal response of trees to drought - a link to vegetation dynamics?, Journal of Vegetation Science, vol.58, issue.3, pp.442-454, 2009.
DOI : 10.1111/j.1654-1103.2009.05701.x